The Principles of Biology, Volume 1 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 1 (of 2)
Spencer, Herbert
Biology
in a reproductive organ--either an ovarium or the homologue of one--has
been called, by Prof. Owen, _parthenogenesis_. It is the process familiarly
exemplified in the _Aphides_. Here, from the fertilized eggs laid by
perfect females there grow up imperfect females, in the ovaria of which are
developed ova that though unfertilized, rapidly assume the organization of
other imperfect females, and are born viviparously. From this second
generation of imperfect females, there by-and-by arises, in the same
manner, a third generation of the same kind; and so on for many
generations: the series being thus symbolized by the letters A, B, B, B, B,
B, &c., A. Respecting this kind of heterogenesis it should be added that,
in animals as in plants, the number of generations of sexless individuals
produced before the re-appearance of sexual ones, is indefinite; both in
the sense that in the same species it may go on to a greater or less extent
according to circumstances, and in the sense that among the generations of
individuals proceeding from the same fertilized germ, a recurrence of
sexual individuals takes place earlier in some of the diverging lines of
multiplication than in others. In trees we see that on some branches
flower-bearing axes arise while other branches are still producing only
leaf-bearing axes; and in the successive generations of _Aphides_ a
parallel fact has been observed. Lastly has to be set down that kind of
heterogenesis in which, along with gamogenesis, there occurs a form of
agamogenesis exactly like it, save in the absence of fecundation. This is
called true parthenogenesis--reproduction carried on by virgin mothers
which are in all respects like other mothers. Among silk-worm-moths this
parthenogenesis is exceptional rather than ordinary. Usually the eggs of
these insects are fertilized; but if they are not they are still laid, and
some of them produce larvæ. In certain _Lepidoptera_, however, of the
groups _Psychidæ_ and _Tineidæ_, parthenogenesis appears to be a normal
process--indeed, so far as is known, the only process; for of some species
the males have never been found.
A general conception of the relations among the different modes of Genesis,
thus briefly described, will be best given by the following tabular
statement.
GENESIS is
{ Oviparous
{ or
Homogenesis, which is usually Gamogenesis { Ovo-viviparous
{ or
{ Viviparous
or
{ Gamogenesis
{ alternating
Heterogenesis, which is { with { Parthenogenesis
{ Agamogenesis { or { Internal
{ Metagenesis { or
{ External
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