The Principles of Biology, Volume 1 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 1 (of 2)
Spencer, Herbert
Biology
re-attached, and supplied with nutriment until it assumes a more or less
complete axial structure.
§ 77. As we have lately seen, the essential act in gamogenesis is the union
of two cell-nuclei, produced in the great majority of cases by different
parent organisms. Nearly always the containing cells, often called
_gametes_, are unlike: the sperm-cell being the male product, and the
germ-cell the female. But among some _Protozoa_ and many of the lower
_Algæ_ and _Fungi_, the uniting cells show no differentiation. Sexuality is
only nascent.
There are very many modes and modifications of modes in which these cells
are produced; very many modes and modifications of modes by which they are
brought into contact; and very many modes and modifications of modes by
which the resulting fertilized germs have secured to them the fit
conditions for their development. But passing over these divergent and
re-divergent kinds of sexual multiplication, which it would take too much
space here to specify, the one universal trait is this coalescence of a
detached portion of one organism with a more or less detached portion of
another.
Such simple _Algæ_ as the _Desmidieæ_, which are sometimes called
unicellular plants, show us a coalescence, not of detached portions of two
organisms, but of two entire organisms: the entire contents of the
individuals uniting to form the germ-mass. Where, as among the
_Confervoideæ_, we have aggregated cells whose individualities are scarcely
at all subordinate to that of the aggregate, the gamogenetic act is often
effected by the union "of separate motile protoplasmic masses produced by
the division of the contents of any cell of the aggregate. These
free-swimming masses of protoplasm, which are quite similar to (but
generally smaller than) the agamogenetic 'zoospores' of the same plants,
and to the free-swimming individuals of many _Protophyta_, are apparently
the primitive type of gametes (conjugating cells); but it is noteworthy
that such a gamete nearly always unites with one derived from another cell
or from another individual. The same fact holds with regard to the gametes
of the Protophytes themselves, which are formed in the same way from the
single cell of the mother individual. In the higher types of
_Confervoideæ_, and in _Vaucheria_, we find these equivalent,
free-swimming, gametes replaced by sexually differentiated sperm- and
germ-cells, in some cases arising in different organs set apart for their
production, and essentially representing those found in the higher plants.
Transitional forms, intermediate between these and the cases where
equivalent gametes are formed from any cell of the plant are also known."
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