The Principles of Biology, Volume 1 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 1 (of 2)
Spencer, Herbert
Biology
Heterogenesis prevails among organisms of which the food, though abundant
compared with their expenditure, is dispersed in such a way that it cannot
be appropriated in a wholesale manner. _Protophyta_, subsisting on diffused
gases and decaying organic matter in a state of minute subdivision, and
_Protozoa_, to which food comes in the shape of extremely small floating
particles, are enabled, by their rapid agamogenetic multiplication, to
obtain materials for growth better than they would do did they not thus
continually divide and disperse in pursuit of it. The higher plants, having
for nutriment the carbonic acid of the air and certain mineral components
of the soil, show us modes of multiplication adapted to the fullest
utilization of these substances. A herb with but little power of forming
the woody fibre requisite to make a stem that can support wide-spreading
branches, after producing a few sexless axes produces sexual ones; and
maintains its race better, by the consequent early dispersion of seeds,
than by a further production of sexless axes. But a tree, able to lift its
successive generations of sexless axes high into the air, where each gets
carbonic acid and light almost as freely as if it grew by itself, may with
advantage go on budding-out sexless axes year after year; since it thereby
increases its subsequent power of budding-out sexual axes. Meanwhile it may
advantageously transform into seed-bearers those axes which, in consequence
of their less direct access to materials absorbed by the roots, are failing
in their nutrition; for it thus throws off from a point at which sustenance
is deficient, a migrating group of germs that may find sustenance
elsewhere. The heterogenesis displayed by animals of the Coelenterate type
has evidently a like utility. A polype, feeding on minute annelids and
crustaceans which, flitting through the water, come in contact with its
tentacles, and limited to that quantity of prey which chance brings within
its grasp, buds out young polypes which, either as a colony or as dispersed
individuals, spread their tentacles through a larger space of water than
the parent alone can; and by producing them, the parent better insures the
continuance of its species than it would do if it went on slowly growing
until its nutrition was nearly balanced by its waste, and then multiplied
by gamogenesis. Similarly with the _Aphis_. Living on sap sucked from
tender shoots and leaves, and able thus to take in but a very small
quantity in a given time, this creature's race is more likely to be
preserved by a rapid asexual propagation of small individuals, which
disperse themselves over a wide area of nutrition, than it would be did the
individual growth continue so as to produce large individuals multiplying
sexually. And then when autumnal cold and diminishing supply of sap put a
check to growth, the recurrence of gamogenesis, or production of fertilized
Public-domain text, read in full here on John Shaqi.
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