The Principles of Biology, Volume 1 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 1 (of 2)
Spencer, Herbert
Biology
§ 104. There is a distribution of organisms in Space, and there is a
distribution of organisms in Time. Looking first at their distribution in
Space, we observe in it two different classes of facts. On the one hand,
the plants and animals of each species have their habitats limited by
external conditions: they are necessarily restricted to spaces in which
their vital actions can be performed. On the other hand, the existence of
certain conditions does not determine the presence of organisms that are
fit for them. There are many spaces perfectly adapted for life of a high
order in which only life of a much lower order is found.
While, in the inevitable restriction of organisms to environments with
which their natures correspond we find a _negative_ cause of distribution,
there remains to be found that _positive_ cause whence results the presence
of organisms in some places appropriate to them and their absence from
other places equally appropriate or more appropriate. Let us consider the
phenomena as thus classed.
§ 105. Facts which illustrate the limiting influence of surrounding
conditions are abundant, and familiar to all readers. It will be needful,
however, here to cite a few typical ones of each order.
The confinement of different kinds of plants and different kinds of
animals, to the media for which they are severally adapted, is the broadest
fact of distribution. We have extensive groups of plants that are
respectively sub-aerial and sub-aqueous; and of the sub-aqueous some are
exclusively marine, while others exist only in rivers and lakes. Among
animals we similarly find some classes confined to the air and others to
the water; and of the water-breathers some are restricted to salt water and
others to fresh water. Less conspicuous is the fact that within each of
these contrasted media there are further widespread limitations. In the
sea, certain organisms exist only between certain depths, and others only
between other depths--the limpet and the mussel within the littoral zone,
and numerous kinds at the bottom of the ocean; and on the land, there are
Floras and Faunas peculiar to low regions and others peculiar to high
regions. Next we have the familiar geographical limitations made by
climate. There are temperatures which restrict each kind of organism
between certain isothermal lines, and hygrometric states which prevent the
spread of each kind of organism beyond areas having a certain humidity or a
certain dryness. Besides such general limitations we find much more special
limitations. Some minute vegetal forms occur only in snow. Hot springs have
their peculiar _Infusoria_. The habitats of certain Fungi are mines or
other dark places. And there are creatures unknown beyond the water
contained in particular caves. After these limits to distribution imposed
by physical conditions, come limits imposed by the presence or absence of
other organisms. Obviously, graminivorous animals are confined within
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