The Principles of Biology, Volume 1 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 1 (of 2)
Spencer, Herbert
Biology
Concerning the special aspects of animal-metabolism, we have first to note
those various minor transformations that are auxiliary to the general
transformation by which force is obtained from food. For many of the vital
activities merely subserve the elaboration of materials for activity at
large, and the getting rid of waste products. From blood passing through
the salivary glands is prepared in large quantity a secretion containing
among other matters a nitrogenous ferment, ptyaline, which, mixed with food
during mastication, furthers the change of its starch into sugar. Then in
the stomach come the more or less varying secretions known in combination
as gastric juice. Besides certain salts and hydrochloric acid, this
contains another nitrogenous ferment, pepsin, which is instrumental in
dissolving the proteids swallowed. To these two metabolic products aiding
solution of the various ingested solids, is presently added that product of
metabolism in the pancreas which, added to the chyme, effects certain other
molecular changes--notably that of such amylaceous matters as are yet
unaltered, into saccharine matters to be presently absorbed. And let us
note the significant fact that the preparation of food-materials in the
alimentary canal, again shows us that unstable nitrogenous compounds are
the agents which, while themselves changing, set up changes in the
carbo-hydrates and proteids around: the nitrogen plays the same part here
as elsewhere. It does the like in yet another viscus. Blood which passes
through the spleen on its way to the liver, is exposed to the action of "a
special proteid of the nature of alkali-albumin, holding iron in some way
peculiarly associated with it." Lastly we come to that all-important organ
the liver, at once a factory and a storehouse. Here several metabolisms are
simultaneously carried on. There is that which until recent years was
supposed to be the sole hepatic process--the formation of bile. In some
liver-cells are masses of oil-globules, which seem to imply a carbo-hydrate
metamorphosis. And then, of leading importance, comes the extensive
production of that animal-starch known as glycogen--a substance which, in
each of the cells generating it, is contained in a plexus of protoplasmic
threads: again a nitrogenous body diffused through a mass which is now
formed out of sugar and is now dissolved again into sugar. For it appears
that this soluble form of carbo-hydrate, taken into the liver from the
intestine, is there, when not immediately needed, stored up in the form of
glycogen, ready to be re-dissolved and carried into the system either for
immediate use or for re-deposit as glycogen at the places where it is
presently to be consumed: the great deposit in the liver and the minor
deposits in the muscles being, to use the simile of Prof. Michael Foster,
analogous in their functions to a central bank and branch banks.
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