The Principles of Biology, Volume 1 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 1 (of 2)
Spencer, Herbert
Biology
An instructive parallelism may be noted between these processes carried on
in the animal organism and those carried on in the vegetal organism. For
the carbo-hydrates named, easily made to assume the soluble or the
insoluble form by the addition or subtraction of a molecule of water, and
thus fitted sometimes for distribution and sometimes for accumulation, are
similarly dealt with in the two cases. As the animal-starch, glycogen, is
now stored up in the liver or elsewhere and now changed into glucose to be
transferred, perhaps for consumption and perhaps for re-deposit; so the
vegetal starch, made to alternate between soluble and insoluble states, is
now carried to growing parts where by metabolic change it becomes cellulose
or other component of tissue and now carried to some place where, changed
back into starch, it is laid aside for future use; as it is in the turgid
inside leaves of a cabbage, the root of a turnip, or the swollen
underground stem we know as a potato: the matter which in the animal is
used up in generating movement and heat, being in the plant used up in
generating structures. Nor is the parallelism even now exhausted; for, as
by a plant starch is stored up in each seed for the subsequent use of the
embryo, so in an embryo-animal glycogen is stored up in the developing
muscles for subsequent use in the completion of their structures.
§ 23e. We come now to the supreme and all-pervading metabolism which has
for its effects the conspicuous manifestations of life--the nervous and
muscular activities. Here comes up afresh a question discussed in the
edition of 1864--a question to be reconsidered in the light of recent
knowledge--the question what particular metabolic changes are they by which
in muscle the energy existing under the form of molecular motion is
transformed into the energy manifested as molar motion?
There are two views respecting the nature of this transformation. One is
that the carbo-hydrate present in muscle must, by further metabolism, be
raised into the form of a nitrogenous compound or compounds before it can
be made to undergo that sudden decomposition which initiates muscular
contraction. The other is the view set forth in § 15, and there reinforced
by further illustrations which have occurred to me while preparing this
revised edition--the view that the carbo-hydrate in muscle, everywhere in
contact with unstable nitrogenous substance, is, by the shock of a small
molecular change in this, made to undergo an extensive molecular change,
resulting in the oxidation of its carbon and consequent liberation of much
molecular motion. Both of these are at present only hypotheses, in support
of which respectively the probabilities have to be weighed. Let us compare
them and observe on which side the evidence preponderates.
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