The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
satisfactorily explained, is scarcely to be expected: the circumstances
of each case have to be studied. But it seems not improbable that they
are due to causes of the kind indicated.[36]
§ 235. We have already glanced at clusters of flowers for the purpose
of considering their shapes as clusters. We must now return to them to
observe the modifications undergone by their component flowers. Among
these occur illustrations of great significance.
An example of transition from the radial to the bilateral form in
clustered flowers of the same species, is furnished by the cultivated
_Geraniums_, called by florists _Pelargoniums_. Some of these, bearing
somewhat small terminal clusters of flowers, which are closely packed
together with their faces almost upwards, have radially-symmetrical
flowers. But among other varieties having terminal clusters of which
the members are mutually thrust on one side by crowding, the flowers
depart very considerably from the radial shape towards the bilateral
shape. A like result occurs under like conditions in Rhododendrons
and Azaleas. The _Verbena_, too, furnishes an illustration of radial
flowers rendered slightly two-sided by the slight two-sidedness
of their relations to other flowers in the cluster. And among the
_Cruciferæ_ a kindred case occurs in the cultivated Candytuft.
Evidence of a somewhat different kind is offered us by clustered
flowers in which the peripheral members of the clusters differ from
the central members; and this evidence is especially significant
where we find allied species that do not exhibit the deviation, at
the same time that they do not fulfil the conditions under which it
may be expected. Thus, in _Scabiosa succisa_, Fig. 250, which bears
its numerous small flowers in a hemispherical knob, the component
flowers, similarly circumstanced, are all equal and all radial; but in
_Scabiosa arvensis_, Fig. 251, in which the numerous small flowers form
a flattened disk only the confined central ones are radial: round the
edge the flowers are much larger and conspicuously bilateral.
[Illustration: Figs. 250, 251.]
[Illustration: Fig. 252.]
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