The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
Composite flowers furnish evidence so nearly allied to that which
clustered flowers furnish, that we may fitly glance at them under
the same head. Such a common type of this order as the Sun-flower,
exemplifies the extremely marked difference which arises in many
of these plants between the closely-packed internal florets, each
similarly circumstanced on all sides, and the external florets, not
similarly circumstanced on all sides. In Fig. 253, representing the
inner and outer florets of a Daisy, the contrast is marked between the
small radial corolla of the one and the larger bilateral corolla of the
other. In many cases, however, this contrast is less marked: the inner
florets also having their outward-growing prolongations--a difference
possibly related to some difference in the habits of the insects that
fertilize them. Nevertheless, these composite flowers which have
inner florets with strap-shaped corollas outwardly directed, equally
conform to the general principle; both in the radial arrangement of
the assemblage of florets, and in the bilateral shape of each floret;
which has its parts alike on the two sides of a line passing from the
centre of the assemblage to the circumference. Certain other members
of this order fulfil the law somewhat differently. In _Centaurea_,
for instance, the inner florets are small and vertical in direction,
while the outer florets are large and lateral in direction. And here
may be remarked, in passing, a clear indication of the effect which
great flexibility of the petals has in preventing a flower from
losing its original radiate form; for while in _C. cyanus_, the large
outward-growing florets, having short, stiff divisions of the corolla,
are decidedly bilateral, in _C. scabiosa_, where the divisions of the
corolla are long and flexible, the radial form is scarcely at all
modified. On bearing in mind the probable relations of the forms to
insect-agency, the meaning of this difference will not be difficult to
understand.[38]
§ 236. In extremely-varied ways there are thus re-illustrated among
flowers, the general laws of form which leaves and branches and
entire plants disclose to us. Composed as each cluster of flowers
is of individuals that are originally similar; and composed as each
flower is of homologous foliar organs; we see both that the like
flowers become unlike and the like parts of each flower become unlike,
where the positions involve unlike incidence of forces. The symmetry
remains radial where the conditions are equal all round; shows
deviation towards two-sidedness where there is slight two-sidedness
of conditions; becomes decidedly bilateral where the conditions are
decidedly bilateral; and passes into an unsymmetrical form where the
relations to the environment are unsymmetrical.
CHAPTER XI.
THE SHAPES OF VEGETAL CELLS.
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