The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
§ 245. Among aggregates of the second order, as among aggregates of the
first order, we find that of those possessing any definite shapes the
lowest are spherical or spheroidal. Such are some of the _Radiolaria_,
as _Collozoum inerme_. These bodies which float passively in the sea,
and present in turn all their sides to the same influences, have their
parts disposed with approximate regularity round a centre--approximate,
because in the absence of locomotion a slight irregularity of
growth, almost certain to take place, may cause a fixed attitude and
a resulting deviation from spherical symmetry. The best cases in
illustration of the truth here named, are furnished by rotating and
locomotive organisms respecting which there is a dispute whether they
are animal or vegetal--the _Volvocineæ_. These, already instanced
under the one head in § 218, may here be instanced afresh under the
other. Further, among these secondary aggregates in which the units,
only physically integrated, have not had their individualities merged
into an individuality of a higher order, must be named the compound
_Infusoria_. The cluster of _Vorticellæ_ in Fig. 144, will sufficiently
exemplify them; and the striking resemblance borne by its individuals
to those of a radially-arranged cluster of flowers, will show how,
under analogous conditions, the general principles of morphological
differentiation are similarly illustrated in the two kingdoms.
§ 246. Radial symmetry is usual in low aggregates of the second order
which have their parts sufficiently differentiated and integrated
to give individualities to them as wholes. The _Cœlenterata_
offer numerous examples of this. Solitary polypes--hydroid or
helianthoid--mostly stationary, and when they move, moving with any
side foremost, do not by locomotion subject their bodies to habitual
contrasts of conditions. Seated with their mouths upwards or downwards,
or else at all degrees of inclination, the individuals of a species
taken together, are subject to no mechanical actions affecting some
parts of their discs more than other parts. And this indeterminateness
of attitude similarly prevents their relations to prey from being such
as subject some of their prehensile organs to forces unlike those to
which the rest are subject. The fixed end is differently conditioned
from the free end, and the two are therefore different; but around the
axis running from the fixed to the free end the conditions are alike
in all directions, and the form therefore is radial. Again, among many
of the simple free-swimming _Hydrozoa_, the same general truth is
exemplified under other circumstances. In a common _Medusa_, advancing
through the water by the rhythmical contractions of its disc, the
mechanical reactions are the same on all sides; and as, from accidental
causes, every part of the edge of the disc comes uppermost in its
turn, no part is permanently affected in a different way from the rest.
Hence the radial form continues.
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