The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
§ 282. To put the co-operative actions in their actual order,
would require us to consider them as working on individuals small
modifications that become conspicuous and definite only by inheritance
and gradual increase; but it will aid our comprehension without leading
us into error, if we suppose the whole process resumed in a single
continuously-existing plant.
As the plant erects the integrated series of fronds whose united
parts form its rudimentary axis, the increasing area of frond surface
exposed to the sun’s rays entails an increasing draught upon the
liquids contained in the rudimentary axis. The currents of sap so
produced, once established along certain lines of cells that offer
least resistance, render them by their continuous passage more and
more permeable. This establishment of channels is aided by the wind.
Each bend produced by it while yet the tissue is undifferentiated,
squeezes towards the place of growth and evaporation the liquids
that are passing by osmosis from cell to cell; and when the lines of
movement become defined, each bend helps, by forcing the liquid along
these lines, to remove obstructions and make continuous canals. As fast
as this transfer of sap is facilitated, so fast is the plant enabled
further to raise itself, and add to its assimilating surfaces; and so
fast do the transverse strains, becoming greater, give more efficient
aid. The canals thus formed can be neither in the centre of the
rudimentary axis nor at its surface: for at neither of these places
can the transverse strains produce any considerable compressions.
They must arise along a tract between the outside of the axis and its
core--a tract along which there occur the severest squeezes between the
stretched outer layers and the internal mass. Just that distribution
which we find, is the distribution which these mechanical actions tend
to establish.
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