The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
§ 284. The inferior _Algæ_, along with little unlikeness of parts,
show us little mutual dependence of parts. Having surfaces similarly
circumstanced everywhere, much physiological division of labour
cannot arise; and therefore there cannot be much physiological unity.
Among the superior _Algæ_, however, the differentiation between the
attached part and the free part is accompanied by some integration.
There is evidently a certain transfer of materials, which is doubtless
facilitated by the elongated forms of the cells in the stem, and
probably leads to the formation of dense tissue at the places of
greatest strain, in a way akin to that recently explained in other
cases. And where there is this co-ordination of actions, the parts are
so far mutually dependent that each dies if detached from the other.
That though the organization is so low neither part can reproduce the
other and survive by so doing, is probably due to the circumstance
that neither part contains any considerable stock of untransformed
protoplasm, out of which new tissues may be produced.
Fungi and Lichens present no very significant advances of integration.
We will therefore pass at once to the Archegoniates. In those of them
which, either as single fronds or strings of fronds, spread over
surfaces, and which, rooting themselves as they spread, do not need
that each part should receive aid from remote parts, there is no
developed vascular system serving to facilitate transfer of nutriment:
the parts being little differentiated there is but little integration.
But along with assumption of the upright attitude and the accompanying
specializations, producing vessels for distributing sap and hard tissue
for giving mechanical support, there arises a decided physiological
division of labour; rendering the aërial part dependent on the embedded
part and the embedded part dependent on the aërial part. Here, indeed,
as elsewhere, these concomitant changes are but two aspects of the
same change. Always the gain of power to discharge a special function
involves a loss of power to perform other functions; and always,
therefore, increased mutual dependence constituting physiological
integration, must keep pace with that increased fitting of particular
parts to particular duties which constitutes physiological
differentiation.
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