The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
Did space permit, it would be possible to point out among the
_Protozoa_, various cases analogous to that of the _Arcella_; which may
be described as like a microscopic Limpet, having a sarcode body of
which the upper surface has become horny, while the lower surface with
its protruding pseudopodia, retains the primitive jelly-like character.
That differentiations of this kind have been gradually established
among these minute creatures through the unlike relations of their
parts to the environment, is an inference supported by a form which,
while the rest of the body has a scarcely distinguishable coating,
“agrees with _Arcella_ and _Difflugia_ in having the pseudopodia
protrusible from one extremity only of the body.”
Many parallel specializations of surface among aggregates of the second
order might be instanced from the _Cœlenterata_. In the _Hydra_, the
ectoderm presents over its whole area no conspicuous unlikenesses;
but there usually exist in the hydroid polypes of superior types,
decided contrasts between the higher and lower parts. While the
higher parts retain their original characters, the lower parts excrete
hard outer layers yielding support and protection. Various stages
of the differentiation might be followed. “In _Hydractinia_,” says
Prof. Green, this horny layer “becomes elevated at intervals to form
numerous rough processes or spines, while over the general surface
of the ectoderm its presence is almost imperceptible.” In other
types, as in _Cordylophora_, it spreads part way up the animal’s
sides, ending indefinitely. In _Bimeria_ it “extends itself so as to
enclose the entire body of each polypite, leaving bare only the mouth
and tips of the tentacles.” While in _Campanularia_ it has become a
partially-detached outer cell, into which the creature can retract its
exposed parts.
But it is as needless as it would be wearisome to trace through the
several sub-kingdoms the rise of these multiform contrasts, with the
view of seeking interpretations of them. It will suffice if we take a
few groups of the illustrations furnished by the higher animals.
§ 292. We may begin with those modifications of surface which subserve
respiration. Though we ordinarily think of respiration as the quite
special function of a quite special organ, yet originally it is not
so. Little-developed animals part with their carbonic acid and absorb
oxygen, through the general surface of the body. Even in the lower
types of the higher classes, the general surface of the body aids
largely in aërating the blood; and the parts which discharge the
greater part of this function are substantially nothing more than
slightly altered and extended portions of the skin.
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