The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
Thus, as Prof. Huxley has somewhere pointed out, there is a
substantial parallelism between all the sensory organs in their modes
of development; as there is, too, between their modes of action.
A _vibrissa_ may be taken as their common type. Increased
impressibility by an external stimulus, requires an increased
peripheral expansion of the nervous system on which the stimulus may
fall; and this is secured by an introversion of the integument, forming
a sac on the walls of which a nerve may ramify. That the more extended
sensory area thus constituted may be acted upon, there requires some
apparatus conveying to it from without the appropriate stimulus; and
in the case of the _vibrissa_, this apparatus is the epidermic
growth which, under the form of a hair, protrudes from the sac. And
that the greatest sensitiveness may be obtained, the external action
must be exaggerated or multiplied by the apparatus which conveys it
to the recipient nerve; as, in the case of the _vibrissa_, it is
by the development of a hair into an elastic lever, that transforms
the slight force acting through considerable space on its exposed end,
into a greater force acting through a smaller space at its rooted end.
Similarly with the organs of the higher senses. In a rudimentary eye,
the slightly modified sense cell has but a rudimentary nerve to take
cognizance of the impression; and to concentrate the impression upon
it, there is nothing beyond a thickening of the epidermis into a
lens-shape. But the developed eye shows us a termination of the nerve
greatly expanded and divided to receive the external stimulus. It shows
us an introverted portion of the integument containing the apparatus
by which the external stimulus is conveyed to the recipient nerve. The
structure developed in this sac not only conveys the stimulus, but
also, like its homologue, concentrates it; and in the one case as in
the other, the structure which does this is an epidermic growth from
the bottom of the sac. Even with the ear it is the same. Again we have
an introverted portion of the integument, on the walls of which the
nerve is distributed in the primitive ear. The otolithes contained
in the sac thus formed, are bodies which are set in motion by the
vibrations of the surrounding water, and convey these vibrations in an
exaggerated form to the nerves. And though it is not alleged that these
otolithes are developed from the epidermic lining of the chamber, yet
as, if not so developed, they are concretions from the contents of an
epidermic sac, they must still be regarded as epidermic products.
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