The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
“With regard to your method of explaining the fundamental characters
of ‘Exogens’ and ‘Endogens,’ this of course is part of the same
hypothesis; but I may point out that since Von Mohl and Sanio, between
1855 and 1865, showed (1) that the growth at the stem apex of a
monocotyledon was _not_ endogenous, and (2) that the ‘thickening ring’
near the apex of a dicotyledon was not to be confused, as had been
done up till then, with the ring of _secondary meristem_ or _true
cambium_, which arose lower down, and only in woody or practically
woody stem, the terms ‘Exogen’ and ‘Endogen’ have necessarily fallen
into disuse, since they imply a false conception of what happens. Both
monocotyledons and dicotyledons have a ‘thickening ring,’ which gives
rise to the primary vascular cylinder of the stem. When the stem is
of considerable thickness, as in Palms, &c., it grows by the active
cell-division of its outer layers, so that both classes are ‘exogenous’
in this sense; while the addition of a centrifugal zone of secondary
wood is confined to certain Dicotyledons (Trees, shrubs, &c.).
“The distinction between the embryos, moreover, is not absolute. The
single cotyledon is usually terminal in monocotyledons, but not always
(_Dioscoraceæ_ have lateral cotyledons), but the plumule may push
through it (Grasses) or make its exit sideways (Palms), or be formed at
the side (_Alisma_); and Dicotyledons very similarly.
“The occurrence of completely sheathing leaves in grasses is perhaps
correlated with the absence of cambium, but grasses are an aberrant
type among monocotyledons, and secondary thickening is only found in
very few genera of this class, so that the correlation is, so to speak,
negative and indirect.... It is clear that the greater part of the
discussion will have to be re-written.”
For the reasons assigned in the preface I cannot undertake to re-write
the discussion, as suggested. It must stand for what it is worth. All I
can do is here to include along with it the foregoing criticisms.
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Elsewhere in the archive
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account