The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
chain of individuals arising by gemmation, and continuing permanently
united end to end. Such a chain can arise only on condition that
combination is more advantageous than separation; and for it to be more
advantageous, the anterior members of the series must become adapted to
functions facilitated by their positions, while the posterior members
become adapted to functions which their positions permit. Hence, direct
or indirect equilibration or both, must tend continually to establish
types in which the connected individuals are more and more unlike
one another, at the same time that their several individualities are
more and more disguised by the integration consequent on their mutual
dependence.
Such being the anticipations warranted by the general laws of
evolution, we have now to inquire whether there are any animals which
fulfil them. Very little search suffices; for structures of the kind
to be expected are abundant. In that great division of the animal
kingdom at one time called _Annulosa_, but now grouped into _Annelida_
and _Arthropoda_, we find a variety of types having the looked-for
characters. Let us contemplate some of them.
§ 205. An adult Chætopod is composed of segments which repeat one
another in their details as well as in their general shapes. Dissecting
one of the lower orders, such as is shown in Fig. 160, proves that
the successive segments, besides having like locomotive appendages,
like branchiæ, and sometimes even like pairs of eyes, also have like
internal organs. Each has its enlargement of the alimentary canal;
each its contractile dilatation of the great blood-vessel; each its
portion of the double nervous cord, with ganglia when these exist;
each its branches from the nervous and vascular trunks answering to
those of its neighbours; each its similarly answering set of muscles;
each its pair of openings through the body-wall; and so on throughout,
even to the organs of reproduction. That is to say, every segment is
in great measure a physiological whole--every segment contains most
of the organs essential to individual life and multiplication: such
essential organs as it does not contain, being those which its position
as one in the midst of a chain, prevents it from having or needing. If
we ask what is the meaning of these homologies, no adequate answer is
supplied by any current hypothesis. That this “vegetative repetition”
is carried out to fulfil a predetermined plan, was shown to be quite
an untenable notion (§§ 133, 134). On the one hand, we found nothing
satisfactory in the conception of a Creator who prescribed to himself
a certain unit of composition for all creatures of a particular class,
and then displayed his ingenuity in building up a great variety of
forms without departing from the “archetypal idea.” On the other hand,
examination made it manifest that even were such a conception worthy
of being entertained, it would have to be relinquished; since in each
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