The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
the resulting segments are so many gemmiparously-produced individuals,
is necessarily less obvious.
[Illustration: Figs. 162–165.]
§ 206. Evidence remains which adds very greatly to the weight of
that already assigned. Thus far we have studied only the individual
segmented animal; considering what may be inferred from its mode of
evolution and final organization. We have now to study segmented
animals in general. Comparison of different groups of them and of
kinds within each group, will disclose various phases of progressive
integration of the nature to be anticipated.
[Illustration: Figs. 166–169.]
Among the simpler _Platyhelminthes_, as in some kinds of _Planaria_,
transverse fission occurs. A portion of a _Planaria_ separated by
spontaneous constriction, becomes an independent individual. Sir J.
G. Dalyell found that in some cases numerous fragments artificially
separated, grew into perfect animals.[21] In these creatures which thus
remind us of the lowest _Hydrozoa_ in their powers of agamogenetic
multiplication, the individuals produced one from another do not
continue connected. As the young ones laterally budded-off by the
_Hydra_ separate when complete, so do the young ones longitudinally
budded-off by the _Planaria_. Fig. 166 indicates this. But there
are allied types which show us a more or less persistent union of
homologous parts, or individuals, similarly arising by longitudinal
gemmation.[22] The cestoid _Entozoa_ furnish illustrations. Without
dwelling on the fact that each segment of a _Tænia_, like each separate
_Planaria_, is an independent hermaphrodite; and without specifying the
sundry common structural traits which add probability to the suspicion
that there is some kinship between the individuals of the one order
and the segments of the other; it will suffice to point out that the
two types are so far allied as to demand their union under the same
sub-class title. And recognizing this kinship, we see significance
in the fact that in the one case the longitudinally-produced gemmæ
separate as complete individuals, and in the other continue united
as segments in smaller or larger numbers and for shorter or longer
periods. In _Tænia echinococcus_, represented in Fig. 167, we have
a species in which the number of segments thus united does not
exceed four. In _Echinobothrium typus_ there are eight or ten; and
in cestoids generally they are numerous.[23] A considerable hiatus
occurs between this phase of integration and the next higher phase
which we meet with; but it is not greater than the hiatus between the
types of the _Platyhelminthes_ and the _Chætopoda_, which present
the two phases. Though it is doubtful whether separation of single
segments occurs among the Annelida,[24] yet very often we find strings
of segments, arising by repeated longitudinal budding, which after
reaching certain lengths undergo spontaneous fission: in some cases
doing this so as to form two or more similar strings of segments
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