The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
§ 208. Insects, Arachnids, Crustaceans, and Myriapods, are all members
of that higher division of the _Annulosa_[26] called _Articulata_
or now more generally _Arthropoda_. Though in these creatures the
formation of segments may be interpreted as a disguised gemmation;
and though, in some of them, the number of segments increases by this
modified budding after leaving the egg, as it does among the Annelids;
yet the process is not nearly so dominant: the segments are usually
much less numerous than we find them in the types last considered. In
most cases, too, the segments are in a greater degree differentiated
one from another, at the same time that they are severally more
differentiated within themselves. Nor is there any instance of
spontaneous fission taking place in the series of segments composing
an articulate animal. On the contrary, the integration, always great
enough permanently to unite the segments, is frequently carried so
far as to hide very completely the individualities of some or many of
them; and occasionally, as among the Acari, the consolidation, or the
arrest of segmentation, is so decided as to leave scarcely a trace
of the articulate structure: the type being in these cases indicated
chiefly by the presence of those characteristically-formed limbs, which
give the alternative name _Arthropoda_ to all the higher _Annulosa_.
Omitting the parasitic orders, which, as in other cases, are aberrant
members of their sub-kingdom, comparisons between the different orders
prove that the higher are strongly distinguished from the lower, by
the much greater degree in which the individuality of the tertiary
aggregate dominates over the individualities of those secondary
aggregates called segments or “somites,” of which it is composed. The
successive Figs. 170–176, representing (without their limbs) a Julus,
a Scolopendra, an isopodous Crustacean, and four kinds of decapodous
Crustaceans, ending with a Crab, will convey at a glance an idea of
the way in which that greater size and heterogeneity reached by the
higher types, is accompanied by an integration which, in the extreme
cases, nearly obliterates all traces of composite structure. In the
Crab the posterior segments, usually folded underneath the shell, alone
preserve their primitive distinctness. So completely confluent are the
rest, that it seems absurd to say that a Crab’s carapace is composed of
as many segments as there are pairs of limbs, foot-jaws, and antennæ
attached to it; and were it not that during early stages of the Crab’s
development the segmentation is faintly marked, the assertion might be
considered illegitimate.
[Illustration: Figs. 170–176.]
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