The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
is but one step to the conception of gemmiparously-produced individuals
which severally part one from another as soon as they are formed.
§ 209. We must now return to a junction whence we diverged some time
ago. As before explained under the head of Classification, organisms
do not admit of uniserial arrangement, either in general or in detail;
but everywhere form groups within groups. Hence, having traced the
phases of morphological composition up to the highest forms in any
sub-kingdom, we find ourselves at the extremity of a great branch, from
which there is no access to another great branch, except by going back
to some place of bifurcation low down in the tree.
There exist such similarities of shape and structure between the larval
forms of low Molluscs and those of Annelids and Rotifers, as to show
that there was an early type common to them all; and its probable
characters, suggested by comparison, seem to imply that it had arisen
from some cœlenterate type, intermediate between the _Cnidaria_ and
the _Ctenophora_. But there is this noteworthy difference between the
molluscan larva and the allied larvæ, that it gives origin to only one
animal and not to a group of animals, united or disunited. No true
Mollusc multiplies by gemmation, either continuous or discontinuous;
but the product of every fertilized germ is a single individual.
It is a significant fact that here, where for the first time we have
homogenesis holding throughout an entire sub-kingdom, we have also
throughout an entire sub-kingdom no case in which the organism is
divisible into two, three, or more, like parts. There is neither any
such clustering or branching as a cœlenterate or molluscoid animal
usually displays; nor is there any trace of that segmentation which
characterizes the _Annulosa_. Among these animals in which no single
egg produces several individuals, no individual is separable into
several homologous divisions. This connexion will be seen to have
a probable meaning, on remembering that it is the converse of the
connexion which obtains among the _Annulosa_, considered as a group.
A Mollusc, then, is an aggregate of the second order. Not only in the
adult animal is there no sign of a multiplicity of like parts that
have become obscured by integration; but there is no sign of such
multiplicity in the embryo. And this unity is just as conspicuous in
the lowest Lamellibranch as in the highest Cephalopod.
[Illustration: Figs. 188–190.]
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