The Principles of Biology, Volume 2 (of 2)Spencer, Herbert
Science
The Principles of Biology, Volume 2 (of 2)
Spencer, Herbert
Biology
§ 211. On glancing back at the facts brought together in these two
chapters, we see it to be probable that there has gone on among animals
a process like that which we saw reason to think has gone on among
plants. Minute aggregates of those physiological units which compose
living protoplasm, exist as _Protozoa_: some of them incoherent,
indefinite, and almost homogeneous, and others of them more coherent,
definite, and heterogeneous. By union of these nucleated particles
of sarcode, are produced various indefinite aggregates of the second
order--Sponges, _Polycytharia_, Foraminifers, &c.; in which the
compound individuality is scarcely enough marked to subordinate the
primitive individualities. But in other types, as in _Hydra_, the lives
of the morphological units are in a considerable degree, though not
wholly, merged in the life of the integrated body they form. As the
primary aggregate, when it passes a certain size, undergoes fission or
gemmation; so does the secondary aggregate. And as on the lower stage
so on the higher, we see cases in which the gemmiparously-produced
individuals part as soon as formed, and other cases in which they
continue united, though in great measure independent. This massing
of secondary aggregates into tertiary aggregates, is variously
carried on among the _Hydrozoa_, the _Actinozoa_, the _Polyzoa_,
and the _Tunicata_. In most of the types so produced, the component
individualities are very little subordinated to the individuality of
the composite mass--there is only physical unity and not physiological
unity; but in certain of the oceanic _Hydrozoa_, the individuals are
so far differentiated and combined as very much to mask them. Forms
showing us clearly the transition to well-developed individuals of
the third order, are not to be found. Nevertheless, in the great
sub-kingdom _Annulosa_, there are traits of structure, development,
and mode of multiplication, which go far to show that its members are
such individuals of the third order; and in the relations to external
conditions involved by the mode of union, we find an adequate cause for
that obscuration of the secondary individualities which we must suppose
has taken place. The two other great subdivisions, _Mollusca_ and
_Vertebrata_, between the lower members of which there are suggestive
points of community, present us only with aggregates of the second
order, that have in many cases become very large and very complex.
We find in them no trace of the union of gemmiparously-produced
individuals. Neither the molluscous nor the vertebrate animal shows
the faintest trace of a segmentation affecting the totality of
its structure; and we see good grounds for concluding that such
segmentation as exceptionally occurs in the one and usually occurs in
the other, is superinduced.
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