The Principles of Stratigraphical GeologyMarr, J. E. (John Edward)
Science
The Principles of Stratigraphical Geology
Marr, J. E. (John Edward)
Geology, Stratigraphic
_The Devonian flora and faunas._ The plant remains in the Lower
Palæozoic rocks are few in number. Some undoubted terrestrial plants
have been discovered, but the prevalent flora of lower Palæozoic
times, so far as yet known, was one consisting of Algæ. In Devonian
times we begin to meet with a number of Cryptogams of higher type,
allied to those which form the dominant flora of the succeeding
period. The fauna is in many ways remarkable. The Devonian period has
been termed the period of ganoid fishes, and the remarkable remains,
so graphically described by the late Hugh Miller, are indeed
peculiarly characteristic of Devonian times, but they are largely
though by no means exclusively entombed in rocks of the Old Red
Sandstone type[86]. The Devon type of rock contains a great abundance
and variety of the problematical group, the Stromatoporoids, which
contribute extensively to the formation of many of the limestones,
and although these organisms are not by any means confined to Devonian
strata, their abundance and variety therein might lead one to speak of
the period as that of Stromatoporoids. The remains of corals are very
abundant in the limestones, and, as already stated, frequently give
rise to true reef-masses. The graptolites, as remarked in the previous
chapter, disappear in the rocks of the Devonian period, and as only
one or two fragments have been found, we may assert that the group was
practically extinct at the end of Silurian times, though species of
one genus, _Monograptus_, lingered for a short time in greatly
diminished quantity. The trilobites which played so important a part
amongst the faunas of Lower Palæozoic times still occur fairly
abundantly amongst the rocks of the Devonian system, and there is a
very interesting point to be noticed in connexion with them. They seem
to have become practically extinct in the succeeding Carboniferous
period, where few genera are found, and the decadence of the group
began in Devonian times. In these circumstances it is interesting to
note the tendency displayed by the creatures to possess spiny
coverings. It is true that _Acidaspis_, the most spinose of all
trilobites, is abundant in Ordovician and Silurian strata, and that
other spinose trilobites are found there, but the peculiarity of the
Devonian trilobites is, that genera which were previously smooth, or
rarely possessing one or few spines, are found represented by
extremely spinose species in these beds,--the spines being developed
from all parts of the test, sometimes as a fringe to head or tail,
sometimes as prominent projections from glabella and neck segment, and
frequently in rows down the body segments. Besides _Acidaspis_, we
find spinose species of _Phacops_, _Homalonotus_, _Cyphaspis_,
_Bronteus_ and _Encrinurus_ in Devonian strata, and the occurrence of
these forms is so frequent and world-wide, that one might perhaps
infer with confidence that an unknown fauna containing many spiny
trilobites was of Devonian age.
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