Two species only, the two which I have named, have proved satisfactory
in both directions. Let us watch them at work. They will show us the
principal features of the manner of life led by the whole tribe, for
they occupy the two extremes of the scale of sizes, the Bull
Onthophagus being one of the largest and the Forked Onthophagus one of
the smallest.
We will speak first of the nest. Contrary to my expectation, the
Onthophagi are indifferent nest-builders. With them we find no spheres
rolled joyously in the sunshine, no ovoids manipulated laboriously in
an underground workshop. Their business, that of reducing filth to
dust, appears to give them so much to do that they have no time left
for work demanding prolonged patience. They confine themselves to what
is strictly necessary and most rapidly obtained.
A perpendicular well is dug, a couple of inches deep, cylindrical in
shape and varying in bore according to the size of the well-sinker. The
pit of the Forked Onthophagus has the diameter of a lead-pencil; that
of the Bull Onthophagus is twice the width. Right at the bottom are the
grub’s provisions, plastered against the walls in a tightly-packed
heap. The total lack of free space at the sides of the pile shows how
the provisioning is done. There is not a sign of a niche, of the least
corner that would leave the mother enough liberty of movement to knead
and mould her bun. The material therefore is simply pressed down at the
bottom of the cylindrical sheath, where it takes the shape of a full
thimble.
I dig up some nests of the Forked Onthophagus near the end of July. It
is a crude piece of work, which surprises you by its roughness when you
think of the neat little worker. Wisps of hay, sticking out anyhow,
increase the untidy look of things. The nature of the materials,
supplied this time by the Mule, are partly the cause of this ugly
appearance.
The length of these nests is fourteen millimetres, the width seven.
[43] The upper surface is slightly concave, proving that the pressure
has been exercised by the mother. The lower end is rounded like the
bottom of the well which serves as a mould. I take a needle and with
the point of it I pick the rustic structure to pieces. The mass of
foodstuff occupies the base, forming the lower two-thirds of the
thimble into a compact block; the cell containing the egg is at the
top, under a thin, concave lid.
There is nothing fresh about the work of the Bull Onthophagus, which,
save for being larger, differs in no way from that of the Forked
Onthophagus. I am unacquainted with the insect’s modus operandi. As
regards the inner secrets of nest-building, these dwarfs are as
reticent as their big colleagues. One alone satisfied my curiosity, or
nearly; and then it was not an Onthophagus but a kindred species, the
Yellow-footed Oniticellus (O. flavipes).
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