The experiment attracted me. I was unable to undertake it for lack of
the means whereby to produce a cold, even temperature over a long time.
What should I have obtained if my penury had not made me abandon the
enterprise? A retarding of the progress of the metamorphosis, but
nothing more, apparently. The horn on the corselet would have persisted
in its sterility and, sooner or later, would have disappeared.
I have reasons for my conviction. The abode of the Onthophagus engaged
on his metamorphosis is not deep down; variations of temperature are
easily felt. On the other hand, the seasons are capricious, especially
the spring. Under the skies of Provence, the months of May and June, if
the mistral lend a hand, have periods when the thermometer drops in
such a way as to suggest a return of winter.
To these vicissitudes add the influence of a more northerly climate.
The Onthophagi occupy a wide zone of latitude. Those of the north, less
favoured by the sun than those of the south, might quite possibly have
the date of their transformation postponed by a change in the weather
and consequently be subjected to a lower temperature for several weeks.
This would spin out the work of evolution and give the thoracic armour
time to harden into horn, at rare intervals, as chance may prescribe.
Here and there, then, the requisite condition of a moderate or even low
temperature at the time of the nymphosis actually exists, without the
need of any artificial agency.
Well, what becomes of this surplus time placed at the service of the
organic labour? Does the promised horn ripen? Not a bit of it: it
withers just as it does under the stimulus of a hot sun. In the records
of entomology I find no mention of an Onthophagus carrying a horn upon
his corselet. No one would even have suspected the possibility of such
an armour, if I had not bruited abroad the strange appearance of the
nymph. The influence of climate, therefore, has nothing to do with the
matter.
As we go more deeply into it, the question becomes more complicated.
The horny appendages of the Onthophagus, the Copris, the Minotaurus and
many others are the male’s prerogative; the female is without them or
wears them only on a reduced and very modest scale. We must look upon
these products as personal ornaments rather than as implements of
labour. The male makes himself fine for the pairing; but, with the
exception of the Minotaurus, who pins down the dry pellet that needs
crushing and holds it in position with his trident, I know none that
uses his armour as a tool. Horns and prongs on the forehead, crests and
crescents on the corselet are the male coquette’s jewels and nothing
more. The other sex requires no such baits to attract suitors: its
femininity is enough; and finery is neglected.
Public-domain text, read in full here on John Shaqi.
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