Another important phenomenon of the equifinality of regulation was
discovered by Morgan. A species of the flatworm *Planaria* was found to
restore its totality out of small pieces either by regeneration proper,
if the pieces were fed, or by a sort of rearrangement of material,
on the basis of its harmonious-equipotentiality, if they were kept
fasting. It is important to note that here we see one of the conditions
determining the choice of the way to restoration, as we also do in the
well-known equifinal restitutions of the root in plants, where the
behaviour of the organism depends on the distance of the operation-wound
from the tip.[73] In *Tubularia* the actual stage of restitution that
has been already reached by the stem when the second operation takes
place, may account for the specification of its future organogenesis,
but this is not at all clearly ascertained at present.
[73] The root may be restored by regeneration proper, or by the
production of adventitious roots, or by one of the side-roots changing
its geotropism from horizontal to positive, according to the smaller or
greater distance of the wound from the tip.
*Clavellina* also shows equifinality in its restitution, as has already
been shortly mentioned. The isolated branchial apparatus may restitute
itself by retro-differentiation to an indifferent stage followed by
renovation; or it may regenerate the intestine-sac in the proper way.
Nothing is known here about the conditions, except perhaps that young
individuals seem more apt to follow the first of these two ways, older
ones the second; but there are exceptions to this rule.
The discussion of other instances of equifinality, though important in
themselves, would not disclose anything fundamentally new, and so we may
close the subject with the remark that nothing can show better than the
fact of the equifinality of restitutions how absolutely inadequate all
our scientific conceptions are when confronted with the actual phenomena
of life itself. By analysis we have found differences of potencies,
according as they are simple or complex; by analysis we have found
differences of “systems,” differences of means, and indeed we were glad
to be able to formulate these differences as strictly as possible: but
now we see how, in defiance of our discriminations, one and the same
species of animals behaves now like one sort of our “systems,” and now
like the other; how it uses now one sort of “potencies,” now another.
But even if it is granted that, in the presence of such phenomena of
life, our endeavour seems to be like a child’s play on the shores of the
ocean, I do not see any other way for us to go, so long, at least, as
our goal is human science--that is, a study of facts as demanded by our
mental organisation.
REMARKS ON “RETRO-DIFFERENTIATION”
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