Among animals there is only one well-studied instance of our first type
of adaptive morphological characters. *Salamandra atra*, the black
salamander, a species which only inhabits regions at least two thousand
feet above sea-level, does not bring forth its young until metamorphosis
has taken place. The larvae, however, may be removed from the mother’s
body at an earlier stage and forced to complete their development in
water. Under these circumstances, as was shown in an excellent memoir
by Kammerer,[80] they will change the whole histological type of their
gills and skin in order to meet the new functional conditions. The
change of the conditions of functioning is very severe here, for whereas
the gills had served for nutrition and respiration in the uterus--by
a process of endosmosis--they now serve for respiration only, and, of
course, are surrounded by quite an abnormal chemical medium.
[80] *Arch. Entw. Mech.* 17, 1904.
TRUE FUNCTIONAL ADAPTATION[81]
[81] Roux, *Gesammelte Abhandlungen*, vol. i. 1895; in particular, *Der
Kampf der Teile im Organismus*, Leipzig, 1881.
But all other cases of morphological adaptation among animals, and
several in the vegetable kingdom too, belong to our second group of
these phenomena, which in our analytical discussion we have called
adaptations to functional changes that result from the very nature
of functioning, and which we shall now call by their ordinary name,
“functional adaptation.”
It was Roux who first saw the importance of this kind of organic
regulation and thought it well to give it a distinguishing name. *By
functioning the organisation of organic tissues becomes better adapted
for functioning.* These words describe better than any others what
happens. It is well known that the muscles get stronger and stronger the
more they are used, and that the same holds for glands, for connective
tissue, etc. But in these cases only quantitative changes come into
account. We meet with functional adaptations of a much more complicated
and important kind, when for instance, as shown by Babák,[82] the
intestine of tadpoles changes enormously in length and thickness
according as they receive animal or vegetable food, being nearly twice
as long in the second case. Besides this the so-called mechanical
adaptations are of the greatest interest.
[82] *Arch. Entw. Mech.* 21, 1906. By a very detailed comparative study
Babák was able to prove that it is the plant proteids to which the
effect of vegetable food is chiefly due; thus we have an adaptation
to digestibility. Mechanical circumstances are only of secondary
importance. (See also Yung.)
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