We have not yet considered one feature of all experiments in
hybridisation, which indeed seems to be the most important of all for
the theory of inheritance, if taken together with the fact of the
pureness of the germs. The rule of Mendel always relates to one single
character of the species or varieties concerned in hybridisation, and
if it deals with more than one character, it regards every one of them
separately; indeed, the rule holds for every one of them irrespective
of the others. We cannot study here how this most important fact of
the independence of the single characters of a species with regard to
inheritance leads to the production of new races, by an abnormal mixture
of those characters. We only take advantage of the fact theoretically,
and in doing so, I believe, we can hardly escape the conclusion that
the independence of the single characters in inheritance, taken
together with the pureness of the germ-cells in the most simple form
of hybrids, proves that there occurs in inheritance a sort of handing
over of single and separate morphogenetic agents which relate to the
single morphogenetic characters of the adult. We may use Bateson’s
word “allelomorphs” for these agents, or units, as they may be called,
thereby giving expression to the fact that the single and separate
units, which are handed over in inheritance, correspond to each other in
nearly related species without being the same.
And so we have at least an inkling of what the material continuity of
inheritance is to mean, though, of course, our “single and separate
morphogenetic agents,” or “units” or “allelomorphs” are in themselves
not much more than unknown somethings described by a word; but even then
they are “somethings.”
Besides the researches relating to the rule of Mendel and its
exceptions, founded, that is, upon a study of the “second” generation of
hybrids, there is another important line of research lately inaugurated
by Herbst, which investigates the first generation in hybridisation.
The hybrids themselves are studied with the special purpose of finding
out whether the type of the single hybrid may change according to the
conditions of its development, both outer and inner. The discoveries
thus made may lead some day to a better understanding of the intimate
nature of the “units” concerned in heredity, and perhaps to some
knowledge of the arranging and ruling factor in morphogenesis also.
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