[151] Of course the inheritance of specific values from the results of
fluctuating variations, leading to new averages of variability (see
p. 265), may also be understood in this manner, the conditions of
nourishment acting upon the adult and upon its germs equally well.
[152] *Berichte üb. d. Sitzung. d. Ges. f. Bot.*, Hamburg, 1887, 3 Heft.
Would it not be possible at least that adaptations which last
for thousands of generations or more might in fact change the
adaptive character into a congenital one? Then we not only should
have inheritance of acquired characters, but should have a sort
of explanation at the same time for the remarkable fact that
certain histological structures of a very adapted kind are formed
ontogenetically before any function exists, as is known to be the
case with the structures in the bones of vertebrates, for instance.
Experiments are going on at Paris, and perhaps in other places of
scientific research also, which, it is hoped, will show that animals
reared in absolute darkness for many generations will lose their
perfectly formed eyes, and that animals from the dark with very
rudimentary eyes will be endowed with properly functioning ones, after
they have been reared in the light for generations. Such a result indeed
would account for the many animals, of the most different groups, which
live in dark caves and possess only rudiments of eyes: functional
adaptation is no longer necessary, so-called atrophy by inactivity sets
in, and the results “acquired” by it are inherited.[153]
[153] Quite recently Kammerer (*Arch. Entw. Mech.* 25, 1907, p. 7) has
published very important experiments on the inheritance of “acquired”
modifications with regard to the peculiarities of reproduction in
*Salamandra atra* and *S. maculosa*. It seems rather improbable--though
not absolutely impossible--that the germ cells were directly affected by
the external modifying agent in this case.
But enough of possibilities. Let us be content at present to know at
least a few real instances with regard to the slowness of the process of
what might be said to be “re-adaptation” in some plants. This process
shows us a way by which our problem may some day be solved; it allows
us to introduce inheritance of acquired characters as a legitimate
hypothesis at least, which not only will explain many of the diversities
in systematics historically, but also can be called, though not a *causa
vera*, yet certainly more than a mere fiction.
OTHER PRINCIPLES WANTED
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