We shall try to sketch the outlines of what might properly be called an
*analytical theory of morphogenesis*; that is, to explain the sum of our
knowledge about organic form-production, gained by experiment and by
logical analysis, in the form of a real system, in which each part will
be, or at least will try to be, in its proper place and in relation with
every other part. Our analytical work will give us ample opportunity of
mentioning many important topics of so-called general physiology also,
irrespective of morphogenesis as such. But morphogenesis is always to
be the centre and starting-point of our analysis. As I myself approach
the subject as a zoologist, animal morphogenesis, as before, will be the
principal subject of what is to follow.
2. ANALYTICAL THEORY OF MORPHOGENESIS[24]
[24] Compare my *Analytische Theorie der organischen Entwickelung*,
Leipzig, 1894, and my reviews in *Ergebnisse der Anatomie und
Entwickelungsgeschichte*, vols. viii. xi. xiv., 1899-1905. A shorter
review is given in *Ergebnisse der Physiologie*, vol. v., 1906. The full
literature will be found in these reviews.
α. THE DISTRIBUTION OF MORPHOGENETIC POTENCIES
*Prospective Value and Prospective Potency*
Wilhelm Roux did not fail to see that the questions of the locality and
the time of all morphogenetic differentiations had to be solved first,
before any problem of causality proper could be attacked. From this
point of view he carried out his fundamental experiments.
It is only in terminology that we differ from his views, if we prefer
to call our introductory chapter an analysis of the distribution of
morphogenetic potencies. The result will be of course rather different
from what Roux expected it would be.
Let us begin by laying down two fundamental concepts. Suppose we have
here a definite embryo in a definite state of development, say a
blastula, or a gastrula, or some sort of larva, then we are entitled
to study any special element of any special elementary organ of this
germ with respect to what is actually to develop out of this very
element in the future actual course of this development, whether it be
undisturbed or disturbed in any way; it is, so to say, the actual, *the
real fate* of our element, that we take in account. I have proposed to
call this real fate of each embryonic part in this very definite line
of morphogenesis its *prospective value* (“prospective Bedeutung” in
German). The fundamental question of the first chapter of our analytical
theory of development may now be stated as follows: Is the prospective
value of each part of any state of the morphogenetic line constant,
*i.e.* is it unchangeable, can it be nothing but one; or is it variable,
may it change according to different circumstances?
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account