We first introduce a second concept: the term *prospective potency*
(“prospective Potenz” in German) of each embryonic element. The term
“prospective morphogenetic potency” is to signify the *possible
fate* of each of those elements. With the aid of our two artificial
concepts we are now able to formulate our introductory question thus:
Is the prospective potency of each embryonic part fully given by
its prospective value in a certain definite case; is it, so to say,
identical with it, or does the prospective potency contain more than the
prospective value of an element in a certain case reveals?
We know already from our historical sketch that the latter is true: that
the actual fate of a part need not be identical with its possible fate,
at least in many cases; that the potency of the first four blastomeres
of the egg of the sea-urchin, for instance, has a far wider range than
is shown by what each of them actually performs in even this ontogeny.
There are more morphogenetic possibilities contained in each embryonic
part than are actually realised in a special morphogenetic case.
As the most important special morphogenetic case is, of course, the
so-called “normal” one, we can also express our formula in terms of
special reference to it: there are more morphogenetic possibilities in
each part than the observation of the normal development can reveal.
Thus we have at once justified the application of analytical experiment
to morphogenesis, and have stated its most important results.
As the introductory experiments about “Entwickelungsmechanik” have shown
already that the prospective potency of embryonic parts, at least in
certain cases, *can* exceed their prospective value--that, at least in
certain cases, it can be different from it--the concept of prospective
potency at the very beginning of our studies puts itself in the centre
of analytical interest, leaving to the concept of prospective value the
second place only. For that each embryonic part actually has a certain
prospective value, a specified actual fate in every single case of
ontogeny, is clear from itself and does not affirm more than the reality
of morphogenetic cases in general; but that the prospective value of the
elements may change, that there is a morphogenetic power in them, which
contains more than actuality; in other words, that the term “prospective
potency” has not only a logical but a factual interest: all these points
amount to a statement not only of the most fundamental introductory
results but also of the actual *problems* of the physiology of form.
If at each point of the germ something else *can* be formed than
actually is formed, why then does there happen in each case just what
happens and nothing else? In these words indeed we may state the chief
problem of our science, at least after the fundamental relation of
the superiority of prospective potency to prospective value has been
generally shown.
Public-domain text, read in full here on John Shaqi.
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