But even this terminology would fail to touch precisely the very
centre of facts: it is not only the simplicity or singularity of
their potencies which characterises the rôle of our systems in
morphogenesis,[57] but far more important with respect to the production
of form are two other leading results of the experimental researches.
The proper act to be performed by every element in each actual case is
in fact a single one, but the potency of any element as such consists
in the possibility of many, nay of indefinitely many, single acts:
that then might justify us in speaking of our systems as “indefinite
equipotential,” were it not that another reason makes another title seem
still more preferable. There are indeed indefinite singular potencies at
work in all of our systems during ontogeny: but the sum of what happens
to arise in every case out of the sum of the single acts performed by
all of the single equipotential cells is not merely a sum but a unit;
that is to say, there exists a sort of harmony in every case among the
*real products* of our systems. The term *harmonious-equipotential
system* therefore seems to be the right one to denote them.
[57] The name of singular-equipotential systems might also be applied
to elementary organs, the single potencies of which are awaked to
organogenesis by specific formative stimuli from without; but that is
not the case in the systems studied in this chapter.
We now shall try first to analyse to its very extremes the meaning of
the statement that a morphogenetic system is harmonious-equipotential.
*The “Harmonious-Equipotential System”*
We have an ectoderm of the gastrula of a starfish here before us; we
know that we may cut off any part of it in any direction, and that
nevertheless the differentiation of the ectoderm may go on perfectly
well and result in a typical little embryo, which is only smaller in
its size than it would normally be. It is by studying the formation of
the highly complicated ciliary band, that these phenomena can be most
clearly understood.
Now let us imagine our ectoderm to be a cylinder instead of being
approximately a sphere, and let us imagine the surface of this cylinder
unrolled. It will give us a plane of two definite dimensions, *a* and
*b*. And now we have all the means necessary for the analytical study of
the differentiation of an harmonious-equipotential system.
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