All of you have seen common sea-anemones or sea-roses, and many of you
will also be familiar with the so-called hydroid polyps. *Tubularia*
is one genus of them: it looks like a sea-anemone in miniature placed
on the top of a stem like a flower. It was known already to Allman
that *Tubularia* is able to restore its flower-like head when that
is lost, but this process was taken to be an ordinary regeneration,
until an American zoologist, Miss Bickford, succeeded in showing that
there was no regeneration process at all, in the proper sense of the
word, no budding of the missing part from the wound, but that the new
tubularian head was restored by the combined work of many parts of the
stem. Further analysis then taught us that *Tubularia* indeed is to be
regarded as the perfect type of an harmonious-equipotential system: you
may cut the stem at whatever level you like: a certain length of the
stem will always restore the new head by the co-operation of its parts.
As the point of section is of course absolutely at our choice, it is
clear, without any further discussion, that the prospective value of
each part of the restoring stem is a “function of its position,” that
it varies with its distance from the end of the stem; and so at once
we discover one of the chief characteristics of our systems. But also
the second point which enters into our formula can be demonstrated in
*Tubularia*: the dependence of the fate of every element on the actual
size of the system. You would not be able to demonstrate this on very
long stems, but if you cut out of a *Tubularia* stem pieces which are
less than ten millimetres in length, you will find the absolute size
of the head restored to be in close relation to the length of the stem
piece, and this dependence, of course, includes the second sort of
dependence expressed in our formula.
The figures will serve to show you a little more concretely what has
been described. The head of *Tubularia* consists of a sort of broad base
with a thin proboscis upon it, both bearing a large number of tentacles;
these tentacles are the first things to be seen as primordia (“Anlagen”)
in the process of restitution. You notice two rings of longitudinal
lines inside the stem; the lines will become walls and then will
separate from the stem until they are only connected with it at their
basal ends; the new tentacles are ready as soon as that has happened,
and a process of growth at the end will serve to drive the new head out
of the so-called perisarc or horny skeleton, which surrounds the stem.
By comparing the two figures, 12 *e*, and *g*, you easily find out that
the absolute lengths of the two tentacle rings are very different, and
that both are in proportion[60] to the actual size of the stem (Fig. 12).
Public-domain text, read in full here on John Shaqi.
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