The mouth of the starfish is in the center of the ventral side. A
ventral or /ambulacral groove/ extends through each arm. In these
grooves the ambulacra, or tube-feet, are arranged in rows. The
/ambulacra/ are hollow, cylindrical bodies, each of which has a
sucker at one end, and is connected at the other end with a little
globular body, the /ampulla/. The ventral groove forms a ridge in the
body-cavity. It is called the /ambulacral zone/, and is formed by a
double row of elongated plates, which meet and form a raised line
along the middle of the inside of the arms, and appear like ribs;
between them lie the ampullæ. These plates are called the /ambulacral
ossicles/. Numerous other ossicles extend over the surface, buried
in the integument of the body, making a calcareous network, or
/exoskeleton/. The ossicles are connected by muscles so that the
animal is not rigid, but is able to bend the body and even creep
through comparatively small places. The ossicles are covered with
/spines/ of two kinds. The very minute ones are the /pedicellariæ/,
and have a jointed end which opens and shuts like a pincers. Their
principal use seems to be to remove waste matter or other substances
from the body, keeping it free and clean. The pedicellariæ are
arranged, in some species, in circles around the [pg206] bases
of the spines, and form groups over the dorsal surface. Examining
the backs of different starfishes with a strong magnifying-glass,
one will be surprised to see the varied grouping of the spines and
pedicellariæ. At the extremity of each arm is a red eye-spot, which
is sensitive to light, and a tube-foot, which has no sucker and which
is believed to act both as an olfactory and as a feeling organ. The
madreporic plate lies at the angle of two of the arms or rays. Water
filters through the madreporic plate and passes by the stone-canal to
the ring-canal around the mouth, and thence to the radial tubes, one
of which extends through each arm and is connected by a branch with
each tube-foot. At the base of each tube-foot there is a globular
reservoir, or ampulla. A valve connects the tube-foot and ampulla.
When the ampulla is contracted, water is forced into and distends
the tube-foot; when the ampulla is distended, it withdraws the water
from the tube-foot, thus acting like a suction-bulb. When water
is sent into the tube-feet they are greatly extended and are then
placed against an object. The water being then withdrawn through the
ampullæ, a vacuum is formed, and the suckers are tightly attached.
By the alternating movements of the ambulacra the animal is dragged
slowly along and moves with a gliding motion. It does not deviate
from a straight path, going over elevations or through depressions
without turning aside; its body conforms to the irregularities of
the road and never bridges over spaces. The stomach extends a little
way into the arms, and a short intestine leads to the excretory
opening in the center of the back.
Public-domain text, read in full here on John Shaqi.
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