The next six pairs of appendages are grouped about the mouth. They
are the /mandibles/, the /maxillæ/, and the /maxillipeds/. The
mandibles are at the mouth-opening, and, being heavy and hard, are
adapted to tearing and grinding; they have a jointed attachment,
the /palpus/, whose office is to keep the mandibles clean. The two
pairs of maxillæ are delicate and leaf-like. The three pairs of
maxillipeds grow gradually larger, the last pair being very prominent
and extending over the other mouth-parts. Next come five pairs of
walking-feet. One or more pairs of these feet have pincer-like ends,
or claws. Some species have the claws immensely developed, as in
lobsters. The claws are the /chelæ/, and the feet which bear the
/chelæ/ are termed the /chelipeds/. The rest of the walking-feet
have generally single, hook-like ends, but are variously modified
in different species. The abdominal segments have six pairs of
appendages, also variously modified. The last segment is without
appendages, but often is extended into a tail, [pg259] or
fin-like expansion. The next to the last segment, in many forms, has
appendages modified into swimming-plates, which extend on each side
of the telson, forming a broad, fan-like caudal extremity.
They have, then, to correspond to the twenty segments of the body,
two pairs of sensory, six pairs of mouth-, and five pairs of
walking-appendages attached to the cephalothorax, and six pairs on
the abdomen. The terminal segment, or telson, is without appendages.
The exopodite is present on the maxillipeds, but disappears from the
walking-feet in the higher forms.
In moulting the /Macrura/ split in the longitudinal line down the
back; in the /Brachyura/ the split occurs across the body at the
point between the thorax and the abdomen.
SUBORDER ~MACRURA~
SHRIMPS, PRAWNS, LOBSTERS, CRAWFISH, AND HERMIT-CRABS
The characteristic features of the /Macrura/ are an elongated body
with the abdomen usually extended; a carapace, somewhat cylindrical;
and the last pair of appendages of the abdomen (which are attached to
the next to last segment) united with the last segment, or telson, to
form a powerful caudal fin, used for swimming backward. The creeping
forms in moving walk forward, but swim backward.
FREE-SWIMMING FORMS: SHRIMPS AND PRAWNS
Public-domain text, read in full here on John Shaqi.
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