In these curious animals the posterior part of the body is not
protected by a crustaceous covering, and therefore the animal seeks
protection by inserting its soft and defenseless abdomen into some
hollow object, usually the shell of a gasteropod mollusk, as the
whelk or the periwinkle (/Buccinum/, /Littorina/). The hermit-crabs
do not always use a shell for this purpose, as they are sometimes
found in the tubes of plant-stems or in sponges. Like other
organisms in the animal world, they seem [pg265] well fitted to
make the best of their surroundings, the body becoming modified to
suit the peculiar conditions under which they live. In those which
inhabit shells the abdomen becomes spiral, in conformity to the
convolutions of the shell. All the abdominal appendages are more or
less atrophied; the sixth pair become like hooks, and these fasten
to the columella of the shell, keeping the crab securely attached
to it. In the female some of the abdominal appendages are hair-like
and are adapted to carrying the eggs. The thorax, being protected,
is protruded from the shell at will. The first pair of feet are much
larger than the others, and are provided with claws (chelæ). The
first right foot is usually much larger than the left, and, besides
the usual functions of capturing and crushing prey, the claw, or
hand, serves as an operculum to close the mouth of the shell when the
crab retires completely within it. The small left hand is shaped to
fill out the parts of the opening not covered by the right one, thus
making a close fit. The next two pairs of feet end in simple hooks,
and are used for walking and dragging the crab along when he travels.
These crabs move about very fast, and the houses upon their backs
seem no encumbrance. [pg266]
[Illustration: /Pagurus bernhardus/: /ch/, chela of first right leg;
/l./4, /l./5, fourth and fifth legs; /t/, abdominal terga; /up/, last
pair of appendages, modified to fasten to the columella of the shell
in which the hermit lives.]
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