The digestive tract, beginning at the mouth, opens immediately into a
muscular throat, which is called the /buccal mass/. Within the buccal
mass there is a long, fleshy ribbon that bears upon its upper surface
numerous rows of sharp teeth; these rows of teeth run crossways on
the ribbon, each row containing more or less prominent median teeth,
with several laterals upon each side, and with marginal teeth on each
side of the laterals. This is called the /radula/, and is used for
tearing or rasping the food. When [pg319] not in use it lies coiled
up within the buccal mass. Beyond the buccal mass, food passes into
the esophagus, which widens out into a muscular pouch or stomach;
this may also be provided with chitinous teeth or hard knobs or
plates for the purpose of further triturating the food. Above and
partially surrounding the stomach is a large reddish or brownish
liver, which furnishes to the alimentary canal the digestive juices.
From the stomach the intestine leads to the terminal anal opening.
There are two organs, serving the function of kidneys, placed
dorsally one upon each side of the body (/nephridia/). A heart lies
within the median line of the body, and consists of a ventricle and
two lateral auricles. The vascular system is almost entirely closed;
only in so far as it is partially open does it essentially differ
from that of higher groups; the blood is whitish or light bluish,
and receives its oxygenation in the gills or along the free mantle
surface, through which it courses in exceedingly thin capillaries. In
this manner respiration is carried on to some extent throughout the
whole outer surface of the body. There are two /branchiæ/ (gills).
These have the appearance of small feathers and are placed upon
each side of the animal posteriorly; they project slightly. Spread
over the top of this animal, fitting it perfectly, is a thin fleshy
covering attached to its dorsal portion like a skin, but hanging
free, like a flap, over the sides, the head, and the posterior end
of the animal. Upon the sides it may just cover the tips of the
branchiæ, but in front the head is seen to project from under it.
The loose-hanging portion of this sac-like covering is somewhat
thickened, and perhaps recurved upward upon itself. Upon the edge of
this flap all the way around may occur a row of cilia, or tentacular
processes. This cloak-like covering is called the mantle, and, as
already noted, is the most peculiar and persistent characteristic
of the /Mollusca/. The "cavity" formed between the loose-hanging
portion of the mantle and the sides of the foot is called the /mantle
cavity/, and contains posteriorly the gills, or branchiæ. From the
outer surface of this mantle a calcareous secretion takes place,
forming a shell, which, in our ideal mollusk, will be shaped in
conformity with the surface outlines of the mantle, oblong and convex
above. Over the lower portion of the shell the edge of [pg320] the
mantle recurves.
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