The /Amphineura/, as already observed, approach most closely to the
ideal mollusk just described. They are bilaterally symmetrical.
This fact is so important that it constitutes them a class,
notwithstanding the fact that in respect to the foot (the basis of
division into classes) they would be included with the /Gasteropoda/,
for (barring some exceptional instances) they creep along upon a foot
quite as our ideal mollusk would, and as the /Gasteropoda/ do. The
head carries no tentacles, thus essentially differing in this respect
from the ideal form. The mantle is extended down in front, completely
covering the head. The branchiæ are confined to a few pairs of
/ctenidia/, or plume-like gills, within the mantle groove or cavity,
and are arranged upon each side of the excretory opening like small
feathers.
There are two orders of the /Amphineura/, the /Polyplacophora/ and
the /Aplacophora/ (or /Solenogastres/), the one name meaning "bearing
many plates," and the other "without plates," the word "plate" in
this sense being synonymous with "shell." The shell of the first
order consists of eight calcareous disks arranged in a longitudinal
row along the back or dorsal side of the animal, which overlap like
shingles on a roof and admit of great variation of form in the
various families. [pg322]
It will be observed from the examples given that the shell plates do
not entirely cover the mantle, but that a considerable portion of the
upper mantle surface is left exposed along the side of the animal.
This exposed portion of the mantle is called the /girdle/. It is
always very much thickened and is often exceedingly tough. In some
species it is covered by a great number of calcareous spicules or
granules which are embedded in the leathery mantle tissue. Strangely
enough, the /Chitonidæ/ (a large family of the /Polyplacophora/)
possess a great number of eyes scattered over the shell plates like
minute dots.
[Illustration: /Chiton spinosus/, dorsal view.]
The members of this order are to be found in all parts of the world,
but their greatest development is upon the west coast of America.
In California the student of marine life has a splendid opportunity
to examine the chitons. They are generally to be found adhering
to the under surface of stones about low-tide mark. They are very
sluggish in their movements and may readily be removed from their
resting-places by suddenly thrusting a knife-blade under the foot
and prying them off. When thus detached they have a habit of rolling
up into a ball and so remaining indefinitely; to prevent this they
should be placed at once in a jar of sea-water, where they will
resume their natural position. For preservation as cabinet specimens,
chitons should be tied down between two boards or flat substances,
to hold them firmly extended until dried; otherwise they are apt to
contract out of shape.
[Illustration:/Chiton/, ventral view: /an/, anus; /cten/, ctenidia;
/ft/, foot; /mant/, mantle-edge; /mo/, mouth.]
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