Let us now take a good example of a gasteropod and locate its various
organs; at the same time we may use the occasion to refer to more
important modifications of these organs which will be encountered
later in the various genera. The most available gasteropod on the
east coast of the United States for this purpose, is /Fulgur/, both
on account of its large size and its abundance. /Buccinum/ may be
used if the student is north of Cape Cod and therefore unable to
secure a good living specimen of /Fulgur/; the anatomical differences
between the two are slight.
Note the siphon protruding forward from a notch in the shell. This
consists merely in an elongation of a fold of the mantle, which is
held in a manner to constitute a tube, through which [pg331] the
animal draws in water to supply the gills, lying just back of the
siphon and concealed under the mantle in the cavity already spoken
of as the mantle cavity. With the exception of that portion of the
mantle which constitutes the siphon, no part of its margin, it will
be observed, can be seen, the edge of the shell completely hiding it
from view.
In some genera the siphon is exceedingly long—in fact, longer than
the body of the animal. In these cases the mollusk is generally fond
of burying itself in the sand and maintaining communication with
the world above only by means of this long, fleshy tube. In some
genera the siphon is protected by an elongation of the shell, as in
/Fasciolaria/ (Plate LXXVI). An extreme example of this is given in
the figure on page 343, which represents a species of /Murex/, where
the siphon is not only protected by the shell, but the shell itself
is further protected by the double row of sharp spines with which it
is beset.
[Illustration: /Buccinum undatum/: /si/, siphon.]
There are some genera which possess no siphons at all, in all of
which cases there is no notch in the shell opening, the aperture
being round, as in /Natica/ and in /Littorina/ (pages 368, 371). The
mollusks of this type make a fold in the forward part of the mantle
which serves as a siphon. Such a muscular folding in the mantle would
be technically called a /functional siphon/.
On the posterior dorsal side of the foot is attached a horny plate,
of concentric structure about a central or subcentral point or
nucleus. This is called the /operculum/, and serves to close the
entrance of the shell when the animal is withdrawn, the object no
doubt being protective. The opercula vary to a great extent in the
different genera; in some cases they are calcareous, in others
cartilaginous. When the aperture of the shell is very large it often
happens that the operculum does not entirely close the opening, as
in /Fulgur/, but in /Buccinum/ it is a close fit. The operculum is
entirely wanting in the /Pulmonata/, one of the [pg332] large orders
of the /Gasteropoda/. The figures represent some of the commoner
types of opercula.
Public-domain text, read in full here on John Shaqi.
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