The classification of the /Gasteropoda/ is primarily a division into
three great orders: the /Opisthobranchiata/, the /Prosobranchiata/,
and the /Pulmonata/. The /Pulmonata/, which include the land and
fresh-water gasteropod shells, are characterized by the presence of
a breathing-sac, or lung, instead of gills. They are, therefore,
essentially air-breathers, and would perish if kept too long under
water. The /Opisthobranchiata/ and /Prosobranchiata/ are marine,
and, like all animals which breathe by means of gills, soon die
when removed from the water. We have already noted, however, how
some genera of marine gasteropods that find their station about
high-tide mark are able to live for considerable periods out of the
water, notwithstanding the fact that they are fully equipped with
gills and not possessed of lungs. Indeed, there is one large family
of prosobranchs (the /Cyclostomatidæ/) that has become entirely
terrestrial in habit, its members having lost their gills and
acquired lungs, but their organization otherwise is so essentially
that of the prosobranchs that they have never been considered as
pulmonates.
The main difference between these two orders of marine gasteropods
is that in the prosobranchs the breathing-organs (the gills) are
placed in a position forward of the heart, and the degree of torsion
(page 330) characteristic of this molluscan class has been continued
until the auricle of the heart is in front of the ventricle. There
is always a shell, usually spiral, and, with few exceptions, an
operculum. In the opisthobranchs, on the other hand, the relative
position of the heart and gills is reversed, and they further differ
from the prosobranchs in that the sexes are always united in each
individual. The opisthobranchs are not always provided with a shell.
One division of the order, known as the "nudibranchs," are entirely
naked. These are commonly called the "sea-slugs," and are to be
found crawling about the marine vegetation in shallow water, in
tide-pools, and on the piling of old docks. Curiously enough, the
sea-slugs (see page 354) have no gills at all, but, having lost these
apparently essential organs, are [pg350] enabled to "breathe water"
through their skin. They are wonderfully beautiful creatures, and the
collector must not fail to secure some specimens, put them into a jar
of sea-water, and watch them expand.
There are other more superficial differences between the prosobranchs
and the opisthobranchs, which will enable even a beginner to
distinguish them at a glance. When the latter are possessed of an
external shell, it is bulbous, generally glassy, and with a simple
lip, the aperture extending the entire length of the shell. Again,
the mantle or the propodium of the foot is greatly extended and
usually covers the shell almost wholly.
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