Three kinds of zoöids, polyps, or hydranths—as they are
indiscriminately called—are attached to the stem. Those having
an [pg120] open end and a crown of tentacles are the nutritive
individuals. Small, club-like dilations are immature zoöids. The
/blastostyles/, or reproductive zoöids, are long, cylindrical,
mouthless, and covered. At maturity the cover is ruptured, and the
medusæ have the appearance of a pile of thin saucers attached by the
middle of the convex side. When at length these saucers are set free
as little medusæ, or jellyfishes, the convex side of each saucer,
or swimming-bell, is called the /ex-umbrella/; the concave, under
side, the /subumbrella/. From the center of the subumbrella projects
the /manubrium/, or stomach of the animal. At the free end of the
manubrium is a four-cornered mouth. From the attached end of the
/manubrium/ four tubes or canals diverge, and, extending through the
animal, open into a circular canal which runs around the margin of
the umbrella. When the medusa is as above described, it has reached
the highest point in its development.
When the medusa has matured, it lays eggs, known as /planulæ/. These
are spherical bodies covered with cilia (hairs), by means of which
they swim about for a time; but they finally attach themselves
to some object, there to grow and develop into hydroid colonies.
The cycle of life is thus completed. This process is known as
/alternation of generation/, or /metagenesis/, one life-history
containing two quite different forms of being. The term of life of an
individual is one year, the zoöphyte stage beginning in the autumn
and the medusa stage in the spring.
Some medusæ, besides reproducing by means of eggs, multiply by
budding, small medusæ growing on the manubrium or on the margin of
the umbrella. /Sarsia/ and /Lizzia/ sometimes increase by budding.
The /Hydrozoa/ are not all of the above type. In the sertularians the
zoöids perish on the stem and have no medusa life, their reproductive
element giving rise to the hydroid form without metamorphosis. The
/Trachylinæ/ have no hydroid life, being always free-swimming medusæ;
others, the /Siphonophora/, live a hydroid life which is unattached,
the colony floating on the ocean; the millepores secrete calcareous
skeletons and always remain fixed, reproducing by budding. [pg121]
Hydroids are very abundant, but are comprised in the few groups
mentioned: namely, those which live only in the fixed colonial state;
those which have alternation of generation, being first hydroids
and then swimming-bells, or medusæ; those which live always in the
medusa state, the eggs of the jellyfish developing at once into
other medusæ; and the /Siphonophora/, or those which have a floating
colonial state, the hydroid never being attached, but floating at
large and capable of locomotion, some of the colony having the
function of propulsion.
Public-domain text, read in full here on John Shaqi.
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