Marine biology -- Great Britain; Seashore; Seashore biology
The description just given applies only to the simplest of the sponges,
and we have now to learn that in the higher members of the group the
structure is much more complicated. In these the surface-pores are the
extremities of very narrow tubes which perforate both layers of the
body-wall and then communicate with wider tubes or spaces within, some
of which are lined with the ciliated cells above described. These
spaces, which are sometimes nearly globular in form, and often arranged
in groups with a common cavity, communicate with wider tubes which join
together until, finally, they terminate in a large opening seen on the
exterior of the sponge. Hence it will be seen that the water entering
the minute pores of the surface has to circulate through a complicated
system of channels and spaces, some of which are lined with the ciliated
cells that urge the current onwards before it is expelled through the
large hole. Further, imagine a number of such structures as we have
described growing side by side, their masses coalescing into one whole,
their inner tubes and spaces united into one complex system by numerous
inter-communications, and having several large holes for the exit of the
circulating water, and you then have some idea of the general nature of
many of the more complex sponges to be found on our shores (see fig.
66).
[Illustration: FIG. 67.--HORNY NETWORK OF A SPONGE, MAGNIFIED]
But even this is not all, for as yet we have been regarding the sponges
as consisting of animal matter only, whereas nearly all of them possess
some kind of internal skeleton for the support of the soft, gelatinous
animal substance. The skeleton consists of matter secreted by certain
cells from material in the water and food, and is either horny,
calcareous, or siliceous. The horny skeleton is formed of a network of
fibres of a somewhat silky character, and often, as in the case of the
toilet sponges, highly elastic; but it is sometimes so brittle that the
sponge mass is easily broken when bent. The fibres of this framework
support not only the outer wall of the sponge, but also the walls of all
the internal tubes and spaces, which are often of so soft a nature that
they would collapse without its aid.
The other forms of skeletons consist of minute bodies of carbonate of
lime or of silica, respectively, which assume certain definite shapes,
resembling stars, anchors, hooks, pins, spindles, &c., and are known as
_spicules_. Such spicules are usually present in those sponges that have
horny skeletons, but in others they form the entire skeleton.
Public-domain text, read in full here on John Shaqi.
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