Marine biology -- Great Britain; Seashore; Seashore biology
It will have been noticed from the above short description that the
pteropod is very unlike the typical Cephalophore as outlined in our
general remarks on the group, especially in the symmetrical form of both
body and shell and in the total or almost total absence of the foot; and
this distinction is so marked that the pteropods are often separated
from all the other _Cephalophora_ into a class by themselves, while all
the remainder are placed in a separate extensive class called the
_Gasteropoda_, because they creep on the ventral surface of the body,
the term signifying stomach-footed.
These gasteropods are divided into four orders: the _Nucleobranchiata_,
in which the respiratory and digestive organs form a nucleus on the
posterior part of the back; the Opisthobranchiata, with gills more or
less exposed towards the rear of the body; the _Pulmonifera_, or
lung-breathing order; and the _Prosobranchiata_, in which the gills are
situated in advance of the heart. The third order includes all the land
snails and slugs, and does not therefore fall within the scope of our
work; but the remaining three consist either exclusively or principally
of marine species, and will be dealt with in the order in which they are
named.
The Nucleobranchs are not really gasteropods in the strictest sense of
the term, for they do not creep along by means of their foot, but all
swim freely in the open ocean, always at the surface, and sometimes
adhere to floating weed by means of a sucker. In fact, the foot of these
creatures is greatly modified in accordance with their habits, one part
being often expanded into a ventral swimming fin, and provided with a
sucking-disc for adhesion, and another produced into a posterior fin for
locomotion.
Like the pteropods, the nucleobranchs are purely pelagic, so that we can
hardly expect to meet with a specimen on or near the shore; and thus we
shall content ourselves with a brief notice of their general characters.
The shell is very variable in size and form, and sometimes even entirely
absent. Large-bodied species often possess but a very small shell, while
some are able to entirely retract themselves and close the mouth of the
shell by an operculum. These animals are generally provided with a large
cylindrical proboscis, and the tongue has recurved teeth. The body is
usually very transparent, often so much so that the blood may be seen
circulating within it, and the nervous system is much more perfectly
developed than in the pteropods. The eyes, too, are perfectly formed.
The presence of special breathing organs may seem to be superfluous in
such delicate and soft-bodied creatures as these, for it may be supposed
that all the oxygen required could be absorbed directly from the water
through their soft structures, as is really the case with many aquatic
creatures; and as a matter of fact some of the nucleobranchs possess no
gills, but others have these organs fully formed.
Public-domain text, read in full here on John Shaqi.
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