Marine biology -- Great Britain; Seashore; Seashore biology
We have already referred to the fact that the shells present a number of
perforations on the exterior, in addition to those which pierce the
partitions within, and it is this characteristic which has led to the
application of the name Foraminifera (hole-bearing) to the little beings
we are considering.
[Illustration: FIG. 55.--A SPIRAL FORAMINIFER SHELL]
[Illustration: FIG. 56.--A FORAMINIFER OUT OF ITS SHELL]
The animal inhabiting the shell is exceedingly simple in structure, even
more so than the amœba. It is merely a speck of protoplasm,
exhibiting hardly any differentiation--nothing, in fact, save a
contractile cavity (the _vacuole_), and numerous granules that probably
represent the indigestible fragments of its food.
The protoplasm fills the shell, and also forms a complete gelatinous
covering on the outside, when the animal is alive; and the vacuole and
granules circulate somewhat freely within the semi-solid mass. Further,
the protoplasm itself is highly contractile, as may be proved by
witnessing the rapidity with which the animal can change its form.
When the foraminifer is alive, it floats freely in the sea, with a
comparatively long and slender thread of its substance protruded through
each hole in the shell. These threads correspond exactly in function
with the blunt pseudopodia of the amœba. Should they come in contact
with a particle of suitable food-material, they immediately surround
it, and rapidly retracting, draw the particle to the surface of the
body. The threads then completely envelop the food, coalescing as soon
as they touch, thus bringing it within the animal.
[Illustration: FIG. 57.--THE SAME FORAMINIFER (FIG. 56) AS SEEN WHEN
ALIVE]
[Illustration: FIG. 58.--SECTION OF THE SHELL OF A COMPOUND
FORAMINIFER]
The foraminifer multiplies by fission, or by a process of budding. In
some species the division of the protoplasm is complete, as in the case
of amœbæ, so that each animal has its own shell which encloses a
single chamber, but in most cases the ‘bud’ remains attached to a parent
cell, and develops a shell that is also fixed to the shell of its
progenitor. The younger animal thus produced from the bud gives rise to
another, which develops in the same manner; and this process continues,
the new bud being always produced on the newest end, till, at last, a
kind of colony of protozoons is formed, their shells remaining attached
to one another, thus producing a compound shell, composed of several
chambers, arranged in the form of a line or spiral, and communicating by
means of their perforated partitions. It will now be seen that each
‘cell’ of the compound protozoon feeds not only for itself, but for all
the members of its colony, since the nourishment imbibed by any one is
capable of diffusion into the surrounding chambers, the protoplasm of
the whole forming one continuous mass by means of the perforated
partitions of the complex skeleton.
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