The Story of a Boulder; or, Gleanings from the Note-book of a Field GeologistGeikie, Archibald
Science
The Story of a Boulder; or, Gleanings from the Note-book of a Field Geologist
Geikie, Archibald
Geology; Petrology
The placoid teeth of the carboniferous rocks show the usual forms
of the order. Some of them are sharp and pointed, as those of the
hybodonts; others have a smooth, rounded, or plate-like form, as in the
cestracionts. The latter often show a dark brilliant surface, and might
be readily enough mistaken for well-worn pebbles. In the oblong rounded
teeth of _psammodus_ the surface is densely covered with minute points
like grains of sand, whence the name of the genus. These teeth, when
sliced and viewed under the microscope by transmitted light, exhibit a
complex reticulated internal structure.
Agassiz' second great Order of fishes is named Ganoid, from a Greek
word signifying brightness, in allusion to the brightly enamelled
surface of their dermal covering. They differ from the placoids in
having their outer surface cased in a strong armature of bone, which
is disposed either in the form of large overlapping plates, as among
the strange tortoise-like fishes of the Old Red Sandstone, or as thick
scales, which are either placed at intervals, as along the back and
sides of the sturgeon, or closely imbricated, as in the stony-gar
(_lepidosteus_) of the American rivers. This strong, massive skeleton
constitutes in many genera the sole support of the animal framework,
the inner skeleton being of a gristly cartilaginous kind, like that
of the skate. On this account traces of the vertebral column are by
no means abundant among the older formations. But as the ganoids form
a sort of intermediate link between the placoid or gristly fishes
on the one hand, and the bony fishes on the other, they are found
to present in their different genera examples of both these kinds
of structure. Thus, the skeleton of the sturgeon consists of a firm
cartilage, out of which the vertebræ are moulded, so that this fish
was at one time ranked with the sharks in the cartilaginous tribe of
Cuvier. The skeletons of some of the older ganoids (as _holoptychius_),
on the other hand, manifest such a decidedly osseous structure, with
sometimes so much of a reptilian cast, that the bones were at first
referred to some huge extinct saurians. The head of the ganoid fishes
is encased in a set of large massive plates of bone, and the jaws
are furnished with several rows of small sharp teeth, intermingled
with a less numerous but larger-sized and more formidable kind. The
interior of the mouth likewise displayed in many ancient genera groups
of palatal teeth, so that the dental apparatus of these animals
must have been very complex and complete. The tail in all the older
ganoids was heterocercal, like that of the sharks, the lobes being
not unfrequently densely covered with minute overlapping scales of
bone--a peculiarity which also extended to the fins. But the fins
were sometimes strengthened in another way by having the foremost ray
greatly thickened and enlarged, so as to form a stiff spine like the
ichthyodorulites of the placoids. The whole of the external surface
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