The question of the origin of the feathers can be discussed only from
the speculative point of view, as they are fully developed in the
Archaeopteryx, and there is no approach toward them in any other living
or fossil organism. But a long discussion of the problem has convinced
scientific men that the feathers are evolved from the scales of the
reptile ancestor. The analogy between the shedding of the coat in a
snake and the moulting of a bird is not uninstructive. In both cases the
outer skin or epidermis is shedding an old growth, to be replaced by
a new one. The covering or horny part of the scale and the feather are
alike growths from the epidermis, and the initial stages of the growth
have certain analogies. But beyond this general conviction that the
feather is a development of the scale, we cannot proceed with any
confidence. Nor need we linger in attempting to trace the gradual
modification of the skeleton, owing to the material change in habits.
The horny beak and the reduction of the toes are features we have
already encountered in the reptile, and the modification of the pelvis,
breast-bone, and clavicle are a natural outcome of flight.
In the Chalk period we find a large number of bird remains, of about
thirty different species, and in some respects they resume the story of
the evolution of the bird. They are widely removed from our modern types
of birds, and still have teeth in the jaws. They are of two leading
types, of which the Ichthyornis and Hesperornis are the standard
specimens. The Ichthyornis was a small, tern-like bird with the power of
flight strongly developed, as we may gather from the frame of its wings
and the keel-shaped structure of its breast-bone. Its legs and feet were
small and slender, and its long, slender jaws had about twenty teeth on
each side at the bottom. No modern bird has teeth; though the fact that
in some modern species we find the teeth appearing in a rudimentary
form is another illustration of the law that animals tend to reproduce
ancestral features in their development. A more reptilian character in
the Ichthyornis group is the fact that, unlike any modern bird, but like
their reptile ancestors, they had biconcave vertebrae. The brain was
relatively poor. We are still dealing with a type intermediate in
some respects between the reptile and the modern bird. The gannets,
cormorants, and pelicans are believed to descend from some branch of
this group.
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