In our study of the evolution of the plant, the insect, and the bird we
were seriously thwarted by the circumstance that their frames,
somewhat frail in themselves, were rarely likely to be entombed in good
conditions for preservation. Earlier critics of evolution used, when
they were imperfectly acquainted with the conditions of fossilisation,
to insinuate that this fragmentary nature of the geological record was a
very convenient refuge for the evolutionist who was pressed for positive
evidence. The complaint is no longer found in any serious work. Where
we find excellent conditions for preservation, and animals suitable
for preservation living in the midst of them, the record is quite
satisfactory. We saw how the chalk has yielded remains of sea-urchins
in the actual and gradual process of evolution. Tertiary beds which
represent the muddy bottoms of tranquil lakes are sometimes equally
instructive in their fossils, especially of shell-fish. The Paludina of
a certain Slavonian lake-deposit is a classical example. It changes
so greatly in the successive levels of the deposit that, if the
intermediate forms were not preserved, we should divide it into several
different species. The Planorbis is another well-known example. In this
case we have a species evolving along several distinct lines into forms
which differ remarkably from each other.
The Tertiary mammals, living generally on the land and only coming by
accident into deposits suitable for preservation, cannot be expected to
reveal anything like this sensible advance from form to form. They were,
however, so numerous in the mid-Tertiary, and their bones are so well
calculated to survive when they do fall into suitable conditions, that
we can follow their development much more easily than that of the birds.
We find a number of strange patriarchal beasts entering the scene in the
early Eocene, and spreading into a great variety of forms in the genial
conditions of the Oligocene and Miocene. As some of these forms advance,
we begin to descry in them the features, remote and shadowy at first, of
the horse, the deer, the elephant, the whale, the tiger, and our other
familiar mammals. In some instances we can trace the evolution with a
wonderful fullness, considering the remoteness of the period and
the conditions of preservation. Then, one by one, the abortive, the
inelastic, the ill-fitted types are destroyed by changing conditions or
powerful carnivores, and the field is left to the mammals which filled
it when man in turn began his destructive career.
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