The primitive giraffe appears suddenly in the later Tertiary deposits of
Europe and Asia. The evidence points to an invasion from Africa, and,
as the region of development is unknown and unexplored, the evolution of
the giraffe remains a matter of speculation. Chevrotains flourished in
Europe and North America in the Oligocene, and are still very primitive
in structure, combining features of the hog and the ruminants. Primitive
deer and oxen begin in the Miocene, and seem to have an earlier
representative in certain American animals (Protoceras), of which the
male has a pair of blunt outgrowths between the ears. The first true
deer are hornless (like the primitive muskdeer of Asia to-day), but by
the middle of the Miocene the males have small two-pronged antlers, and
as the period proceeds three or four more prongs are added. It is some
confirmation of the evolutionary embryonic law that we find the antlers
developing in this way in the individual stag to-day. A very
curious race of ruminants in the later Tertiary was a large antelope
(Sivatherium) with four horns. It had not only the dimensions, but
apparently some of the characters, of an elephant.
The elephant itself, the last type of the Ungulates, has a clearer line
of developments. A chance discovery of fossils in the Fayum district in
Egypt led Dr. C. W. Andrews to make a special exploration, and on the
remains which he found he has constructed a remarkable story of the
evolution of the elephant. [*] It is clear that the elephant was developed
in Africa, and a sufficiently complete series of remains has been found
to give a good idea of the origin of its most distinctive features.
In the Eocene period there lived in the Egyptian region an animal,
something like the tapir in size and appearance, which had its second
incisors developed into small tusks and--to judge from the nasal opening
in the skull--a somewhat prolonged snout. This animal (Moeritherium)
only differed from the ordinary primitive Ungulate in these incipient
elephantine features. In the later Eocene a larger and more advanced
animal, the Palaeomastodon, makes its appearance. Its tusks are larger
(five or six inches long), its molars more elephantine, the air-cells
at the back of the head more developed. It would look like a small
elephant, except that it had a long snout, instead of a flexible trunk,
and a projecting lower jaw on which the snout rested.
*See this short account, "Guide to the Elephants in the
British Museum," 1908.
Public-domain text, read in full here on John Shaqi.
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