The Rodents are represented in primitive form early in the Eocene
period. The teeth are just beginning to show the characteristic
modification for gnawing. A large branch of the family, the Tillodonts,
attained some importance a little later. They are described as combining
the head and claws of a bear with the teeth of a rodent and the general
characters of an ungulate. In the Oligocene we find primitive squirrels,
beavers, rabbits, and mice. The Insectivores also developed some of the
present types at an early date, and have since proved so unprogressive
that some regard them as the stock from which all the placental mammals
have arisen.
The Cetacea (whales, porpoises, etc.) are already represented in the
Eocene by a primitive whale-like animal (Zeuglodon) of unknown origin.
Some specimens of it are seventy feet in length. It has large teeth,
sometimes six inches long, and is clearly a terrestrial mammal that
has returned to the waters. Some forms even of the modern whale develop
rudimentary teeth, and in all forms the bony structure of the fore limbs
and degenerate relic of a pelvis and back limbs plainly tell of the
terrestrial origin. Dolphins appear in the Miocene.
Finally, the Edentates (sloths, anteaters, and armadilloes) are
represented in a very primitive form in the early Eocene. They are then
barely distinguishable from the Condylarthra and Creodonta, and seem
only recently to have issued from a common ancestor with those groups.
In the course of the Tertiary we find them--especially in South America,
which was cut off from the North and its invading Carnivores during
the Eocene and Miocene--developed into large sloths, armadilloes, and
anteaters. The reconnection with North America in the Pliocene allowed
the northern animals to descend, but gigantic sloths (Megatherium) and
armadilloes (Glyptodon) flourished long afterwards in South America.
The Megatherium attained a length of eighteen feet in one specimen
discovered, and the Glyptodon often had a dorsal shield (like that of
the armadillo) from six to eight feet long, and, in addition, a stoutly
armoured tail several feet long.
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