First, the plain issue between the Mendelians and the other two
schools--whether the passage from species to species is brought about
by a series of small variations during a long period or by a few large
variations (or "mutations") in a short period--is open to an obvious
compromise. It is quite possible that both views are correct, in
different cases, and quite impossible to find the proportion of each
class of cases. We shall see later that in certain instances where the
conditions of preservation were good we can sometimes trace a perfectly
gradual advance from species to species. Several shellfish have been
traced in this way, and a sea-urchin in the chalk has been followed,
quite gradually, from one end of a genus to the other. It is significant
that the advance of research is multiplying these cases. There is no
reason why we may not assume most of the changes of species we have
yet seen to have occurred in this way. In fact, in some of the lower
branches of the animal world (Radiolaria, Sponges, etc.) there is often
no sharp division of species at all, but a gradual series of living
varieties.
On the other hand we know many instances of very considerable sudden
changes. The cases quoted by Mendelists generally belong to the plant
world, but instances are not unknown in the animal world. A shrimp
(Artemia) was made to undergo considerable modification, by altering the
proportion of salt in the water in which it was kept. Butterflies have
been made to produce young quite different from their normal young by
subjecting them to abnormal temperature, electric currents, and so on;
and, as I said, the most remarkable effects have been produced on eggs
and embryos by altering the chemical and physical conditions. Rats--I
was informed by the engineer in charge of the refrigerating room on
an Australian liner--very quickly became adapted to the freezing
temperature by developing long hair. All that we have seen of the past
changes in the environment of animals makes it probable that these
larger variations often occur. I would conclude, therefore, that
evolution has proceeded continuously (though by no means universally)
through the ages, but there were at times periods of more acute change
with correspondingly larger changes in the animal and plant worlds.
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