The general characteristic of these early Amphibia is that they very
clearly retain the marks of their fish ancestry. All of them have tails;
all of them have either scales or (like many of the fishes) plates of
bone protecting the body. In some of the younger specimens the gills can
still be clearly traced, but no doubt they were mainly lung-animals. We
have seen how the fish obtained its lungs, and need add only that this
change in the method of obtaining oxygen for the blood involved certain
further changes of a very important nature. Following the fossil
record, we do not observe the changes which are taking place in the
soft internal organs, but we must not lose sight of them. The heart, for
instance, which began as a simple muscular expansion or distension of
one of the blood-vessels of some primitive worm, then doubled and
became a two-chambered pump in the fish, now develops a partition in
the auricle (upper chamber), so that the aerated blood is to some extent
separated from the venous blood. This approach toward the warm-blooded
type begins in the "mud-fish," and is connected with the development
of the lungs. Corresponding changes take place in the arteries, and we
shall find that this change in structure is of very great importance in
the evolution of the higher types of land-life. The heart of the higher
land-animals, we may add, passes through these stages in its embryonic
development.
Externally the chief change in the Amphibian is the appearance of
definite legs. The broad paddle of the fin is now useless, and its main
stem is converted into a jointed, bony limb, with a five-toed foot,
spreading into a paddle, at the end. But the legs are still feeble,
sprawling supports, letting the heavy body down almost to the ground.
The Amphibian is an imperfect, but necessary, stage in evolution. It is
an improvement on the Dipneust fish, which now begins to dwindle very
considerably in the geological record, but it is itself doomed to give
way speedily before one of its more advanced descendants, the Reptile.
Probably the giant salamander of modern Japan affords the best
suggestion of the large and primitive salamanders of the Coal-forest,
while the Caecilia--snake-like Amphibia with scaly skins, which live
underground in South America--may not impossibly be degenerate survivors
of the curious Aistopods.
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