From the evolutionary point of view these new classes are as obscure in
their origin, yet as manifestly undergoing evolution when they do
fully appear, as the earlier classes we have considered. All are of a
primitive and generalised character; that is to say, characters
which are to-day distributed among widely different groups were then
concentrated and mingled in one common ancestor, out of which the later
groups will develop. All belong to the lowest orders of their class. No
Hymenopters (ants, bees, and wasps) or Coleopters (beetles) are found
in the Coal-forest; and it will be many millions of years before the
graceful butterfly enlivens the landscapes of the earth. The early
insects nearly all belong to the lower orders of the Orthopters
(cockroaches, crickets, locusts, etc.) and Neuropters (dragon-flies,
may-flies, etc.). A few traces of Hemipters (now mainly represented by
the degenerate bugs) are found, but nine-tenths of the Carboniferous
insects belong to the lowest orders of their class, the Orthopters and
Neuropters. In fact, they are such primitive and generalised insects,
and so frequently mingle the characteristics of the two orders, that
one of the highest authorities, Scudder, groups them in a special
and extinct order, the Palmodictyoptera; though this view is not now
generally adopted. We shall find the higher orders of insects making
their appearance in succession as the story proceeds.
Thus far, then, the insects of the Coal-forest are in entire harmony
with the principle of evolution, but when we try to trace their origin
and earlier relations our task is beset with difficulties. It goes
without saying that such delicate frames as those of the earlier insects
had very little chance of being preserved in the rocks until the special
conditions of the forest-age set in. We are, therefore, quite prepared
to hear that the geologist cannot give us the slenderest information.
He finds the wing of what he calls "the primitive bug" (Protocimex),
an Hemipterous insect, in the later Ordovician, and the wing of a
"primitive cockroach" (Palaeoblattina) in the Silurian. From these we
can merely conclude that insects were already numerous and varied. But
we have already, in similar difficulties, received assistance from the
science of zoology, and we now obtain from that science a most important
clue to the evolution of the insect.
Public-domain text, read in full here on John Shaqi.
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