In the wide, open forest glades, tree-ferns almost precisely similar to those of the modern tropics reared their leafy crowns. But among them was one peculiar type, in which the fronds were borne in pairs on opposite sides of the stem, leaving when they fell two rows of large horseshoe-shaped scars marking the sides of the trunk. Botanists, who have been puzzled with these plants almost as much as with the Stigmaria, have supposed these scars to be marks of branches, of cones, and even of aërial roots; but specimens in my collection prove conclusively that the stem of this genus was a great caudex made up of the bases of two rows of huge leaves cemented together probably by intervening cellular tissue. As in the Devonian and in modern times, the stems of the tree-ferns of the Carboniferous strengthened themselves by immense bundles of cord-like aërial roots, which look like enormous fossil brooms, and are known under the name Psaronius. We have only time to glance at the vast brakes of tall Calamites which fringe the Sigillaria woods, and stretch far sea-ward over tidal flats. They were allied to modern Mares' Tails or Equisetums, but were of gigantic size, and much more woody structure of stem. The Calamites grew on wet mud and sand-flats, and also in swamps; and they appear to have been especially adapted to take root in and clothe and mat together soft sludgy material recently deposited or in process of deposition. When the seed or spore of a Calamite had taken root, it probably produced a little low whorl of leaves surrounding one small joint, from which another and another, widening in size, arose, producing a cylindrical stem, tapering to a point below. To strengthen the unstable base, the lower joints, especially if the mud had been accumulating around the plant, shot out long roots instead of leaves, while secondary stems grew out of the sides at the surface of the soil, and in time there was a stool of Calamites, with tufts of long roots stretching downwards, like an immense brush, into the mud. When Calamites thus grew on inundated flats, they would, by causing the water to stagnate, promote the elevation of the surface by new deposits, so that their stems gradually became buried; but this only favoured their growth, for they continually pushed out new stems, while the old buried ones shot out bundles of roots instead of regular whorls of leaves.